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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression (NX) levels for human tissues was created by combining the data from these three transcriptomics datasets (HPA, GTEx and FANTOM5).
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Antibody staining in the annotated cell types in the current human tissue is reported as not detected, low, medium, or high. This score is based on the staining intensity and fraction of stained cells.
The RNA-seq details section shows detailed information about the individual samples used for the transcript profiling and results of the RNA-seq analysis.
Information about each individual sample is listed below, including gender, age, a tissue section image and estimated fractions of cell types. pTPM (transcripts per million) values give a quantification of the gene abundance which is comparable between different genes and samples.
Cells in seminiferous ducts: 65 Other cell types: 35
Male, age 34
Testis sample 278 pTPM: 946.3
Cell types%
Cells in seminiferous ducts: 75 Other cell types: 25
Male, age 62
Testis sample 279 pTPM: 624.6
Cell types%
Cells in seminiferous ducts: 70 Other cell types: 30
Male, age 34
Testis sample 280 pTPM: 293.4
Cell types%
Cells in seminiferous ducts: 60 Other cell types: 40
Male, age 26
Testis sample 281 pTPM: 1401.5
Cell types%
Cells in seminiferous ducts: 55 Other cell types: 45
Male, age 50
Testis sample 282 pTPM: 1229.3
Cell types%
Cells in seminiferous ducts: 85 Other cell types: 15
Male, age 37
Testis sample 283 pTPM: 902.6
Cell types%
Cells in seminiferous ducts: 35 Other cell types: 65
Male, age 51
Testis sample 349 pTPM: 2040.3
Cell types%
Cells in seminiferous ducts: 65 Other cell types: 35
Male, age 34
Testis sample 354 pTPM: 1186.2
Cell types%
Cells in seminiferous ducts: 70 Other cell types: 30
Male, age 26
Testis sample 371 pTPM: 1988.4
Cell types%
Cells in seminiferous ducts: 75 Other cell types: 25
TESTIS - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as average pTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. pTPM values of the individual samples are presented next to the box plot.
Max pTPM: 2037.4
Min pTPM: 0.2
Std pTPM: 385.9
Median pTPM: 861.5
GTEX-1A3MX-1026-SM-731F8
60-69 years, male
2037.4
GTEX-17HII-0126-SM-7KFSS
50-59 years, male
1712.1
GTEX-11NV4-1726-SM-5N9FC
60-69 years, male
1683.1
GTEX-12WSH-0326-SM-5GCNH
50-59 years, male
1610.2
GTEX-1GMRU-0626-SM-7MKH2
50-59 years, male
1601.9
GTEX-S7PM-0626-SM-4AD4Q
60-69 years, male
1540.7
GTEX-11ZUS-2726-SM-5FQUA
30-39 years, male
1537.4
GTEX-T6MN-2026-SM-4DM7L
50-59 years, male
1476.0
GTEX-11EQ8-1426-SM-5EGJR
60-69 years, male
1451.3
GTEX-17HHY-0726-SM-7IGM6
50-59 years, male
1428.2
GTEX-1B8SF-2926-SM-731DL
60-69 years, male
1425.5
GTEX-1C64N-2826-SM-7IGP5
50-59 years, male
1418.9
GTEX-11WQC-2326-SM-5EQKE
60-69 years, male
1414.2
GTEX-15G19-1326-SM-6LPIR
60-69 years, male
1410.3
GTEX-1C6VR-1526-SM-7MKG9
60-69 years, male
1379.2
GTEX-14ABY-0626-SM-5Q5C9
50-59 years, male
1362.7
GTEX-WK11-0326-SM-4OOS6
50-59 years, male
1358.6
GTEX-18A66-0726-SM-72D77
60-69 years, male
1352.4
GTEX-13FHP-2826-SM-5IJFW
50-59 years, male
1347.1
GTEX-144GL-0726-SM-5LU4P
50-59 years, male
1345.6
GTEX-183FY-1926-SM-7KFRI
20-29 years, male
1331.5
GTEX-145MO-0126-SM-5S2QU
60-69 years, male
1294.7
GTEX-Y111-2426-SM-4TT23
50-59 years, male
1289.0
GTEX-111VG-1926-SM-5GIDO
60-69 years, male
1286.5
GTEX-U8T8-1126-SM-4DXUE
60-69 years, male
1285.5
GTEX-13OVL-0426-SM-5IFG6
50-59 years, male
1276.0
GTEX-QEG4-0126-SM-48TZE
30-39 years, male
1275.8
GTEX-1C4CL-1926-SM-731DY
30-39 years, male
1266.8
GTEX-117YW-1526-SM-5EGGP
50-59 years, male
1266.7
GTEX-WHSB-2126-SM-4M1XF
50-59 years, male
1252.7
GTEX-QEG5-0126-SM-4R1JR
20-29 years, male
1247.4
GTEX-11EQ9-1926-SM-5PNVV
30-39 years, male
1245.3
GTEX-ZTSS-1526-SM-51MTC
40-49 years, male
1243.6
GTEX-ZYT6-2726-SM-5GICP
30-39 years, male
1240.9
GTEX-WHSE-0426-SM-4M1XO
20-29 years, male
1239.0
GTEX-13112-0226-SM-5P9IV
50-59 years, male
1236.0
GTEX-13N1W-2626-SM-5IJEP
70-79 years, male
1228.8
GTEX-117XS-2026-SM-5GID1
60-69 years, male
1209.9
GTEX-11GS4-2026-SM-5N9CP
60-69 years, male
1203.0
GTEX-12ZZY-0126-SM-5LZV2
60-69 years, male
1201.2
GTEX-17MF6-2726-SM-7IGMZ
40-49 years, male
1200.4
GTEX-XAJ8-1326-SM-47JYT
40-49 years, male
1197.8
GTEX-14BIL-2226-SM-73KWF
50-59 years, male
1190.3
GTEX-13OW5-2526-SM-5L3I1
60-69 years, male
1188.9
GTEX-1EU9M-1626-SM-7EWF2
60-69 years, male
1186.6
GTEX-OOBK-2126-SM-3LK5T
40-49 years, male
1174.0
GTEX-13NZA-2526-SM-5IJFX
60-69 years, male
1173.8
GTEX-SNOS-1126-SM-4DM67
40-49 years, male
1172.4
GTEX-14PJ2-2026-SM-6AJAQ
50-59 years, male
1171.3
GTEX-139TS-1726-SM-5IJG5
60-69 years, male
1166.2
GTEX-ZUA1-2726-SM-59HLJ
40-49 years, male
1164.2
GTEX-REY6-0126-SM-48FDT
60-69 years, male
1163.3
GTEX-ZAB5-2426-SM-5CVMW
50-59 years, male
1163.0
GTEX-12WSL-2326-SM-5DUXQ
50-59 years, male
1160.0
GTEX-14E1K-1926-SM-73KWS
50-59 years, male
1158.1
GTEX-Q2AI-1226-SM-48U14
40-49 years, male
1150.0
GTEX-N7MS-0126-SM-3TW8O
60-69 years, male
1145.2
GTEX-OXRL-2126-SM-3NM98
50-59 years, male
1129.8
GTEX-1B996-2326-SM-731EC
60-69 years, male
1128.1
GTEX-11DXZ-2126-SM-59881
50-59 years, male
1127.3
GTEX-ZPU1-2126-SM-57WED
40-49 years, male
1122.4
GTEX-TKQ2-1526-SM-4DXUN
50-59 years, male
1111.9
GTEX-11OF3-1826-SM-5987N
60-69 years, male
1102.7
GTEX-S4Q7-1226-SM-4AD5I
20-29 years, male
1098.7
GTEX-14DAR-1126-SM-793AT
50-59 years, male
1094.0
GTEX-12WSI-2126-SM-5GCMV
50-59 years, male
1089.0
GTEX-13VXU-0726-SM-5J2O7
50-59 years, male
1087.6
GTEX-144GM-0426-SM-5Q5C8
20-29 years, male
1086.3
GTEX-V1D1-2126-SM-4JBH4
50-59 years, male
1078.1
GTEX-1EKGG-2726-SM-79OOD
60-69 years, male
1073.0
GTEX-17KNJ-1126-SM-7KFT6
50-59 years, male
1072.3
GTEX-Z93S-1726-SM-5HL8G
50-59 years, male
1064.4
GTEX-16YQH-1826-SM-6LPJW
30-39 years, male
1064.3
GTEX-1AX9I-1826-SM-72D5I
60-69 years, male
1058.4
GTEX-18A6Q-2626-SM-7KFT3
60-69 years, male
1057.0
GTEX-13111-1526-SM-5EGJX
50-59 years, male
1051.3
GTEX-OHPM-2126-SM-3LK75
50-59 years, male
1050.6
GTEX-1212Z-0326-SM-5FQSJ
60-69 years, male
1038.5
GTEX-139TT-2226-SM-5LZWO
60-69 years, male
1034.2
GTEX-111FC-1926-SM-5GZYC
60-69 years, male
1033.4
GTEX-131XE-0426-SM-5IJF4
50-59 years, male
1028.6
GTEX-14C38-0126-SM-5YY9V
50-59 years, male
1021.2
GTEX-1CB4G-2426-SM-7DUGM
60-69 years, male
1017.7
GTEX-ZVTK-0126-SM-57WDG
20-29 years, male
1008.6
GTEX-ZA64-1626-SM-5CVME
20-29 years, male
1003.4
GTEX-11TUW-2226-SM-5EQL9
60-69 years, male
1002.2
GTEX-PW2O-1426-SM-48TCD
20-29 years, male
1001.6
GTEX-YEC3-1726-SM-5IFIK
50-59 years, male
1000.2
GTEX-ZZ64-1126-SM-5GZXY
20-29 years, male
995.2
GTEX-U4B1-1526-SM-4DXSL
40-49 years, male
980.2
GTEX-12C56-1426-SM-5FQSW
20-29 years, male
979.5
GTEX-17MFQ-0826-SM-793C8
20-29 years, male
977.1
GTEX-1A3MV-0726-SM-72D5M
30-39 years, male
972.4
GTEX-ZQUD-2026-SM-51MSM
30-39 years, male
968.7
GTEX-15SHV-1926-SM-6LPI1
60-69 years, male
968.3
GTEX-1F6IF-0526-SM-7MKHD
60-69 years, male
966.3
GTEX-OIZH-2126-SM-3NB1P
50-59 years, male
959.7
GTEX-UPJH-0126-SM-4IHLL
50-59 years, male
952.1
GTEX-13NZB-2026-SM-5MR4M
30-39 years, male
945.5
GTEX-13O21-1226-SM-5J2MK
50-59 years, male
940.0
GTEX-12696-0226-SM-5EGL3
60-69 years, male
938.9
GTEX-11O72-0726-SM-5P9GO
40-49 years, male
936.7
GTEX-T8EM-0126-SM-4DM5R
40-49 years, male
936.4
GTEX-X5EB-2026-SM-4E3KA
40-49 years, male
935.3
GTEX-OIZI-0126-SM-3NB13
40-49 years, male
934.2
GTEX-14753-0626-SM-5Q5CY
60-69 years, male
933.8
GTEX-18A7B-1526-SM-7KFTH
70-79 years, male
932.3
GTEX-15RIF-1826-SM-6M469
20-29 years, male
927.3
GTEX-R55D-0126-SM-48FEL
50-59 years, male
927.2
GTEX-XBEC-0126-SM-4GIDT
50-59 years, male
920.4
GTEX-16AAH-1626-SM-7EWE2
50-59 years, male
919.9
GTEX-17GQL-1726-SM-718B8
60-69 years, male
915.3
GTEX-132QS-1226-SM-5P9GD
60-69 years, male
910.7
GTEX-13FLW-2126-SM-5N9FD
50-59 years, male
907.8
GTEX-14C39-0526-SM-664OF
40-49 years, male
906.1
GTEX-SNMC-1026-SM-4DM7K
20-29 years, male
902.5
GTEX-18D9B-2726-SM-72D73
60-69 years, male
902.2
GTEX-18465-1526-SM-7KFTV
50-59 years, male
897.4
GTEX-16Z82-2726-SM-7KULV
60-69 years, male
894.0
GTEX-U8XE-0126-SM-4E3I3
30-39 years, male
891.6
GTEX-ZDYS-1326-SM-5IJFF
50-59 years, male
880.4
GTEX-11NUK-2626-SM-5A5MB
50-59 years, male
877.9
GTEX-XQ3S-2726-SM-4BOP2
20-29 years, male
877.9
GTEX-T5JC-0726-SM-4DM55
20-29 years, male
873.7
GTEX-ZV7C-2026-SM-5NQ8F
50-59 years, male
873.5
GTEX-16MTA-1126-SM-6LPJA
50-59 years, male
872.0
GTEX-TKQ1-0926-SM-4DXU2
30-39 years, male
871.6
GTEX-15RIE-0626-SM-6M47G
60-69 years, male
867.5
GTEX-WYJK-1826-SM-4ONDM
50-59 years, male
866.4
GTEX-XLM4-1526-SM-4AT6D
60-69 years, male
861.5
GTEX-1C64O-2126-SM-7IGPB
60-69 years, male
861.3
GTEX-13OVK-2226-SM-6LPJY
40-49 years, male
856.8
GTEX-QV31-1126-SM-4R1K4
30-39 years, male
845.6
GTEX-18QFQ-1926-SM-72D5H
30-39 years, male
842.0
GTEX-13N2G-0126-SM-5N9DV
60-69 years, male
840.2
GTEX-XGQ4-2026-SM-4AT6G
50-59 years, male
833.8
GTEX-V955-1926-SM-4KL1L
60-69 years, male
828.4
GTEX-17F9E-1626-SM-7EWDB
50-59 years, male
825.8
GTEX-WH7G-1926-SM-4LVMM
40-49 years, male
814.6
GTEX-S95S-1126-SM-4B64E
60-69 years, male
810.5
GTEX-1B8KE-1426-SM-7EWEL
50-59 years, male
798.8
GTEX-OOBJ-2126-SM-3NB1N
60-69 years, male
792.3
GTEX-WOFM-1126-SM-4OOSB
20-29 years, male
791.4
GTEX-111YS-2026-SM-5EGGL
60-69 years, male
790.6
GTEX-R55E-0726-SM-48FCZ
20-29 years, male
788.4
GTEX-ZVZP-2226-SM-57WBF
50-59 years, male
786.6
GTEX-14PHX-1426-SM-69LPP
50-59 years, male
786.5
GTEX-X3Y1-2626-SM-4PQZI
40-49 years, male
781.0
GTEX-QLQW-1026-SM-447A9
30-39 years, male
780.4
GTEX-RUSQ-2126-SM-47JXK
50-59 years, male
776.7
GTEX-13O61-2026-SM-5J2M6
60-69 years, male
774.1
GTEX-1AX9J-1226-SM-72D6P
60-69 years, male
761.6
GTEX-13O1R-0726-SM-5IJEI
60-69 years, male
760.9
GTEX-1339X-1926-SM-5PNVP
40-49 years, male
760.0
GTEX-139T6-1226-SM-5IFFC
50-59 years, male
759.2
GTEX-XBED-2026-SM-4AT5D
60-69 years, male
752.0
GTEX-11EI6-2226-SM-5EGJM
60-69 years, male
746.1
GTEX-S3XE-1526-SM-4AD5A
50-59 years, male
744.8
GTEX-Q2AH-1526-SM-48TZG
40-49 years, male
740.5
GTEX-ZT9W-2226-SM-57WFU
50-59 years, male
739.5
GTEX-17HGU-0726-SM-7DUFN
50-59 years, male
739.2
GTEX-S7SE-0326-SM-4AT5Q
50-59 years, male
738.3
GTEX-WVLH-2626-SM-4MVNV
50-59 years, male
737.0
GTEX-R55C-1426-SM-48FED
40-49 years, male
730.0
GTEX-Y5V6-1726-SM-4VDSZ
60-69 years, male
728.9
GTEX-11P82-1526-SM-5BC5M
20-29 years, male
723.6
GTEX-11NSD-1026-SM-5N9BE
20-29 years, male
722.7
GTEX-X261-2326-SM-4PQYU
50-59 years, male
721.1
GTEX-WZTO-0326-SM-4PQYZ
40-49 years, male
713.8
GTEX-14E7W-0726-SM-664OK
70-79 years, male
710.0
GTEX-14PJ3-1626-SM-664O5
50-59 years, male
703.3
GTEX-XPT6-1626-SM-4B655
20-29 years, male
702.6
GTEX-YF7O-2026-SM-4W1YE
50-59 years, male
693.3
GTEX-U3ZM-1626-SM-4DXSK
40-49 years, male
692.1
GTEX-16XZZ-0626-SM-6M47T
60-69 years, male
689.9
GTEX-147JS-0126-SM-5S2TW
60-69 years, male
687.9
GTEX-11TT1-2226-SM-5GU6B
20-29 years, male
687.7
GTEX-RWSA-2426-SM-47JXR
40-49 years, male
684.7
GTEX-144GN-1626-SM-5Q5BU
50-59 years, male
681.9
GTEX-XMK1-2026-SM-4B65K
40-49 years, male
677.1
GTEX-QV44-1726-SM-4R1KG
50-59 years, male
665.1
GTEX-11LCK-2326-SM-5HL53
30-39 years, male
662.8
GTEX-14E6E-1026-SM-664N9
30-39 years, male
661.2
GTEX-11P7K-1026-SM-5HL6Y
30-39 years, male
646.4
GTEX-111CU-1726-SM-5EGHM
50-59 years, male
644.9
GTEX-P4QS-2126-SM-3NMCF
60-69 years, male
644.1
GTEX-12WSM-1326-SM-5GCP9
60-69 years, male
632.6
GTEX-1BAJH-0726-SM-7IGMF
70-79 years, male
623.6
GTEX-13NYB-2226-SM-5MR58
40-49 years, male
611.1
GTEX-17HHE-1226-SM-793C6
50-59 years, male
609.9
GTEX-1E1VI-1026-SM-7MKGQ
50-59 years, male
597.9
GTEX-ZT9X-1426-SM-5DUX1
40-49 years, male
589.8
GTEX-VJYA-1426-SM-4KL1Y
60-69 years, male
587.6
GTEX-YEC4-1526-SM-4W1YU
40-49 years, male
585.1
GTEX-Y8E4-2226-SM-5LU94
40-49 years, male
573.3
GTEX-14C5O-2326-SM-73KYU
60-69 years, male
562.1
GTEX-14PJ4-1726-SM-664OC
60-69 years, male
560.2
GTEX-NFK9-0126-SM-3LK5H
40-49 years, male
554.4
GTEX-15RJ7-1826-SM-7KUMJ
40-49 years, male
553.4
GTEX-YB5E-1926-SM-5IFIG
40-49 years, male
547.0
GTEX-WFG8-1926-SM-4LVM1
20-29 years, male
533.0
GTEX-14B4R-1026-SM-5TDDS
30-39 years, male
529.0
GTEX-QMRM-1526-SM-4R1K6
50-59 years, male
523.0
GTEX-RM2N-1326-SM-48FCW
50-59 years, male
513.2
GTEX-147GR-0626-SM-5S2PK
60-69 years, male
508.9
GTEX-145MH-2326-SM-5O9AW
50-59 years, male
504.2
GTEX-13FTW-1326-SM-5LZZD
40-49 years, male
498.7
GTEX-1AX9K-1826-SM-731CY
50-59 years, male
483.5
GTEX-QLQ7-1426-SM-4R1JX
60-69 years, male
479.4
GTEX-YJ89-0626-SM-4TT3Z
60-69 years, male
476.4
GTEX-13OVH-0726-SM-5N9BU
70-79 years, male
471.9
GTEX-RN64-2326-SM-48FDW
50-59 years, male
471.8
GTEX-145LT-0426-SM-5LUAP
40-49 years, male
471.2
GTEX-1399R-1626-SM-5P9GG
30-39 years, male
470.6
GTEX-Y9LG-1726-SM-4VBQE
30-39 years, male
457.0
GTEX-ZYFC-0126-SM-5GIEH
50-59 years, male
454.3
GTEX-XPVG-2226-SM-4B65U
50-59 years, male
449.8
GTEX-1CAMR-1526-SM-79OLP
20-29 years, male
449.5
GTEX-WFON-2026-SM-4LVMW
40-49 years, male
433.0
GTEX-U3ZH-1526-SM-4DXV1
30-39 years, male
428.4
GTEX-YFCO-1726-SM-4W21S
40-49 years, male
426.9
GTEX-147F4-0626-SM-5LUAK
50-59 years, male
422.2
GTEX-11ONC-2226-SM-5HL6D
60-69 years, male
411.3
GTEX-S33H-0126-SM-4AD62
30-39 years, male
401.4
GTEX-14PJN-1626-SM-68727
30-39 years, male
397.4
GTEX-ZAB4-0126-SM-5CVMG
40-49 years, male
385.7
GTEX-ZTX8-1126-SM-51MRM
20-29 years, male
370.7
GTEX-Y3I4-2026-SM-4TT6Z
50-59 years, male
334.4
GTEX-O5YT-2126-SM-3MJGD
20-29 years, male
316.6
GTEX-1CB4I-0926-SM-7MKFY
50-59 years, male
313.8
GTEX-13OW6-0126-SM-5IJGM
50-59 years, male
294.5
GTEX-18A67-1826-SM-7KFT7
50-59 years, male
272.9
GTEX-1399T-1526-SM-5P9J6
60-69 years, male
270.4
GTEX-14A5H-0626-SM-5TDCO
60-69 years, male
269.1
GTEX-15SKB-0326-SM-6M477
60-69 years, male
266.6
GTEX-14A6H-2326-SM-5Q5B5
30-39 years, male
265.7
GTEX-PLZ6-1226-SM-3P5ZS
30-39 years, male
241.5
GTEX-PLZ5-1526-SM-3P5ZX
50-59 years, male
236.1
GTEX-1AYCT-1026-SM-79ONR
50-59 years, male
155.2
GTEX-1E2YA-2826-SM-7EPIN
50-59 years, male
151.6
GTEX-145MF-1726-SM-5LU9H
50-59 years, male
140.2
GTEX-ZDTT-2026-SM-5K7TY
60-69 years, male
123.7
GTEX-ZLFU-2026-SM-4WWG2
40-49 years, male
88.9
GTEX-12BJ1-1326-SM-5BC5P
60-69 years, male
68.8
GTEX-117YX-2026-SM-5GIEF
50-59 years, male
50.6
GTEX-11DXY-0226-SM-5H123
60-69 years, male
43.0
GTEX-13OW8-0526-SM-5KM24
60-69 years, male
30.7
GTEX-13QJ3-0226-SM-5S2PU
50-59 years, male
26.4
GTEX-1B8KZ-1826-SM-73KVJ
50-59 years, male
26.0
GTEX-1AX8Z-2826-SM-73KTZ
60-69 years, male
22.7
GTEX-S4Z8-2126-SM-4AD5H
60-69 years, male
21.2
GTEX-1C6VS-1126-SM-7EWEW
60-69 years, male
19.9
GTEX-WY7C-2226-SM-4ONCS
50-59 years, male
11.5
GTEX-17F97-2726-SM-7IGQ4
50-59 years, male
1.3
GTEX-NPJ8-1226-SM-3MJHM
40-49 years, male
1.1
GTEX-1C6VQ-1426-SM-73KTW
50-59 years, male
1.0
GTEX-148VJ-2526-SM-5TDEI
70-79 years, male
0.9
GTEX-WVJS-2726-SM-4MVNU
50-59 years, male
0.3
GTEX-1CB4J-1626-SM-79OOK
60-69 years, male
0.2
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TESTIS - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.