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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression (NX) levels for human tissues was created by combining the data from these three transcriptomics datasets (HPA, GTEx and FANTOM5).
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Antibody staining in the annotated cell types in the current human tissue is reported as not detected, low, medium, or high. This score is based on the staining intensity and fraction of stained cells.
The RNA-seq details section shows detailed information about the individual samples used for the transcript profiling and results of the RNA-seq analysis.
Information about each individual sample is listed below, including gender, age, a tissue section image and estimated fractions of cell types. pTPM (transcripts per million) values give a quantification of the gene abundance which is comparable between different genes and samples.
Cells in seminiferous ducts: 65 Other cell types: 35
Male, age 34
Testis sample 278 pTPM: 860.3
Cell types%
Cells in seminiferous ducts: 75 Other cell types: 25
Male, age 62
Testis sample 279 pTPM: 903
Cell types%
Cells in seminiferous ducts: 70 Other cell types: 30
Male, age 34
Testis sample 280 pTPM: 1014.3
Cell types%
Cells in seminiferous ducts: 60 Other cell types: 40
Male, age 26
Testis sample 281 pTPM: 838.6
Cell types%
Cells in seminiferous ducts: 55 Other cell types: 45
Male, age 50
Testis sample 282 pTPM: 849.7
Cell types%
Cells in seminiferous ducts: 85 Other cell types: 15
Male, age 37
Testis sample 283 pTPM: 970.5
Cell types%
Cells in seminiferous ducts: 35 Other cell types: 65
Male, age 51
Testis sample 349 pTPM: 473.6
Cell types%
Cells in seminiferous ducts: 65 Other cell types: 35
Male, age 34
Testis sample 354 pTPM: 923.6
Cell types%
Cells in seminiferous ducts: 70 Other cell types: 30
Male, age 26
Testis sample 371 pTPM: 669.6
Cell types%
Cells in seminiferous ducts: 75 Other cell types: 25
TESTIS - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as average pTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. pTPM values of the individual samples are presented next to the box plot.
Max pTPM: 2166.8
Min pTPM: 642.3
Std pTPM: 171.7
Median pTPM: 951.8
GTEX-NPJ8-1226-SM-3MJHM
40-49 years, male
2166.8
GTEX-1C6VQ-1426-SM-73KTW
50-59 years, male
1657.3
GTEX-17F97-2726-SM-7IGQ4
50-59 years, male
1432.7
GTEX-1399T-1526-SM-5P9J6
60-69 years, male
1423.2
GTEX-PLZ6-1226-SM-3P5ZS
30-39 years, male
1422.7
GTEX-11DXY-0226-SM-5H123
60-69 years, male
1417.5
GTEX-18A67-1826-SM-7KFT7
50-59 years, male
1383.7
GTEX-1CB4J-1626-SM-79OOK
60-69 years, male
1366.0
GTEX-1E2YA-2826-SM-7EPIN
50-59 years, male
1351.2
GTEX-148VJ-2526-SM-5TDEI
70-79 years, male
1335.6
GTEX-Y3I4-2026-SM-4TT6Z
50-59 years, male
1291.8
GTEX-YJ89-0626-SM-4TT3Z
60-69 years, male
1287.0
GTEX-18465-1526-SM-7KFTV
50-59 years, male
1281.6
GTEX-XBED-2026-SM-4AT5D
60-69 years, male
1273.8
GTEX-1C6VS-1126-SM-7EWEW
60-69 years, male
1230.7
GTEX-QEG5-0126-SM-4R1JR
20-29 years, male
1227.5
GTEX-1AX8Z-2826-SM-73KTZ
60-69 years, male
1222.4
GTEX-ZLFU-2026-SM-4WWG2
40-49 years, male
1213.3
GTEX-NFK9-0126-SM-3LK5H
40-49 years, male
1212.8
GTEX-13QJ3-0226-SM-5S2PU
50-59 years, male
1193.0
GTEX-1A3MX-1026-SM-731F8
60-69 years, male
1184.8
GTEX-PW2O-1426-SM-48TCD
20-29 years, male
1182.6
GTEX-S4Z8-2126-SM-4AD5H
60-69 years, male
1177.6
GTEX-145LT-0426-SM-5LUAP
40-49 years, male
1174.6
GTEX-147F4-0626-SM-5LUAK
50-59 years, male
1170.3
GTEX-S3XE-1526-SM-4AD5A
50-59 years, male
1162.3
GTEX-1B8KZ-1826-SM-73KVJ
50-59 years, male
1159.6
GTEX-P4QS-2126-SM-3NMCF
60-69 years, male
1156.7
GTEX-1399R-1626-SM-5P9GG
30-39 years, male
1156.1
GTEX-V955-1926-SM-4KL1L
60-69 years, male
1153.9
GTEX-147GR-0626-SM-5S2PK
60-69 years, male
1151.5
GTEX-OOBJ-2126-SM-3NB1N
60-69 years, male
1142.2
GTEX-X3Y1-2626-SM-4PQZI
40-49 years, male
1139.4
GTEX-T5JC-0726-SM-4DM55
20-29 years, male
1136.6
GTEX-18A7B-1526-SM-7KFTH
70-79 years, male
1132.9
GTEX-1AYCT-1026-SM-79ONR
50-59 years, male
1132.1
GTEX-17HII-0126-SM-7KFSS
50-59 years, male
1130.8
GTEX-WY7C-2226-SM-4ONCS
50-59 years, male
1115.9
GTEX-17KNJ-1126-SM-7KFT6
50-59 years, male
1114.2
GTEX-U8T8-1126-SM-4DXUE
60-69 years, male
1111.5
GTEX-YEC4-1526-SM-4W1YU
40-49 years, male
1104.4
GTEX-WYJK-1826-SM-4ONDM
50-59 years, male
1097.5
GTEX-16YQH-1826-SM-6LPJW
30-39 years, male
1095.5
GTEX-XLM4-1526-SM-4AT6D
60-69 years, male
1095.2
GTEX-183FY-1926-SM-7KFRI
20-29 years, male
1086.7
GTEX-YF7O-2026-SM-4W1YE
50-59 years, male
1085.8
GTEX-1F6IF-0526-SM-7MKHD
60-69 years, male
1082.6
GTEX-U3ZH-1526-SM-4DXV1
30-39 years, male
1080.6
GTEX-QV31-1126-SM-4R1K4
30-39 years, male
1078.3
GTEX-13N2G-0126-SM-5N9DV
60-69 years, male
1073.6
GTEX-14A6H-2326-SM-5Q5B5
30-39 years, male
1069.6
GTEX-11TT1-2226-SM-5GU6B
20-29 years, male
1064.7
GTEX-14C38-0126-SM-5YY9V
50-59 years, male
1062.4
GTEX-WVLH-2626-SM-4MVNV
50-59 years, male
1058.7
GTEX-QLQW-1026-SM-447A9
30-39 years, male
1052.2
GTEX-12WSM-1326-SM-5GCP9
60-69 years, male
1051.3
GTEX-1GMRU-0626-SM-7MKH2
50-59 years, male
1050.2
GTEX-WFG8-1926-SM-4LVM1
20-29 years, male
1049.2
GTEX-ZV7C-2026-SM-5NQ8F
50-59 years, male
1049.0
GTEX-1BAJH-0726-SM-7IGMF
70-79 years, male
1043.1
GTEX-T8EM-0126-SM-4DM5R
40-49 years, male
1042.6
GTEX-ZT9X-1426-SM-5DUX1
40-49 years, male
1042.5
GTEX-QV44-1726-SM-4R1KG
50-59 years, male
1037.1
GTEX-S33H-0126-SM-4AD62
30-39 years, male
1036.2
GTEX-TKQ1-0926-SM-4DXU2
30-39 years, male
1035.5
GTEX-11O72-0726-SM-5P9GO
40-49 years, male
1033.0
GTEX-17HHY-0726-SM-7IGM6
50-59 years, male
1029.7
GTEX-17HGU-0726-SM-7DUFN
50-59 years, male
1029.6
GTEX-XGQ4-2026-SM-4AT6G
50-59 years, male
1029.2
GTEX-S7SE-0326-SM-4AT5Q
50-59 years, male
1026.6
GTEX-145MO-0126-SM-5S2QU
60-69 years, male
1026.1
GTEX-13OW6-0126-SM-5IJGM
50-59 years, male
1025.4
GTEX-OIZH-2126-SM-3NB1P
50-59 years, male
1022.8
GTEX-R55E-0726-SM-48FCZ
20-29 years, male
1022.7
GTEX-R55C-1426-SM-48FED
40-49 years, male
1018.8
GTEX-15SKB-0326-SM-6M477
60-69 years, male
1017.8
GTEX-18A6Q-2626-SM-7KFT3
60-69 years, male
1012.1
GTEX-15RJ7-1826-SM-7KUMJ
40-49 years, male
1011.8
GTEX-17MF6-2726-SM-7IGMZ
40-49 years, male
1010.7
GTEX-ZQUD-2026-SM-51MSM
30-39 years, male
1010.4
GTEX-WHSE-0426-SM-4M1XO
20-29 years, male
1010.0
GTEX-Q2AH-1526-SM-48TZG
40-49 years, male
1008.7
GTEX-WOFM-1126-SM-4OOSB
20-29 years, male
1006.7
GTEX-14PJ4-1726-SM-664OC
60-69 years, male
1001.3
GTEX-13OW8-0526-SM-5KM24
60-69 years, male
1001.0
GTEX-U3ZM-1626-SM-4DXSK
40-49 years, male
1000.5
GTEX-ZAB4-0126-SM-5CVMG
40-49 years, male
1000.1
GTEX-XPVG-2226-SM-4B65U
50-59 years, male
1000.0
GTEX-13NYB-2226-SM-5MR58
40-49 years, male
999.8
GTEX-17GQL-1726-SM-718B8
60-69 years, male
999.0
GTEX-144GL-0726-SM-5LU4P
50-59 years, male
997.8
GTEX-REY6-0126-SM-48FDT
60-69 years, male
995.7
GTEX-139TS-1726-SM-5IJG5
60-69 years, male
995.5
GTEX-RM2N-1326-SM-48FCW
50-59 years, male
995.3
GTEX-1AX9K-1826-SM-731CY
50-59 years, male
995.0
GTEX-11EI6-2226-SM-5EGJM
60-69 years, male
993.8
GTEX-T6MN-2026-SM-4DM7L
50-59 years, male
992.9
GTEX-1339X-1926-SM-5PNVP
40-49 years, male
989.8
GTEX-TKQ2-1526-SM-4DXUN
50-59 years, male
988.5
GTEX-ZA64-1626-SM-5CVME
20-29 years, male
985.8
GTEX-14E6E-1026-SM-664N9
30-39 years, male
983.3
GTEX-1C4CL-1926-SM-731DY
30-39 years, male
982.3
GTEX-WVJS-2726-SM-4MVNU
50-59 years, male
981.9
GTEX-ZDTT-2026-SM-5K7TY
60-69 years, male
981.8
GTEX-11GS4-2026-SM-5N9CP
60-69 years, male
978.0
GTEX-QMRM-1526-SM-4R1K6
50-59 years, male
977.0
GTEX-1B8SF-2926-SM-731DL
60-69 years, male
976.9
GTEX-15G19-1326-SM-6LPIR
60-69 years, male
976.8
GTEX-13112-0226-SM-5P9IV
50-59 years, male
974.4
GTEX-1B996-2326-SM-731EC
60-69 years, male
971.5
GTEX-ZYFC-0126-SM-5GIEH
50-59 years, male
971.3
GTEX-139TT-2226-SM-5LZWO
60-69 years, male
970.8
GTEX-XQ3S-2726-SM-4BOP2
20-29 years, male
970.6
GTEX-18D9B-2726-SM-72D73
60-69 years, male
970.0
GTEX-11ONC-2226-SM-5HL6D
60-69 years, male
969.2
GTEX-1C64O-2126-SM-7IGPB
60-69 years, male
965.7
GTEX-14ABY-0626-SM-5Q5C9
50-59 years, male
963.6
GTEX-11EQ8-1426-SM-5EGJR
60-69 years, male
962.5
GTEX-S95S-1126-SM-4B64E
60-69 years, male
961.3
GTEX-XPT6-1626-SM-4B655
20-29 years, male
961.2
GTEX-R55D-0126-SM-48FEL
50-59 years, male
960.3
GTEX-UPJH-0126-SM-4IHLL
50-59 years, male
960.2
GTEX-SNOS-1126-SM-4DM67
40-49 years, male
959.5
GTEX-SNMC-1026-SM-4DM7K
20-29 years, male
957.3
GTEX-15RIE-0626-SM-6M47G
60-69 years, male
956.7
GTEX-13N1W-2626-SM-5IJEP
70-79 years, male
956.4
GTEX-144GN-1626-SM-5Q5BU
50-59 years, male
956.1
GTEX-YFCO-1726-SM-4W21S
40-49 years, male
953.2
GTEX-15SHV-1926-SM-6LPI1
60-69 years, male
952.8
GTEX-Y9LG-1726-SM-4VBQE
30-39 years, male
951.8
GTEX-17HHE-1226-SM-793C6
50-59 years, male
951.6
GTEX-WHSB-2126-SM-4M1XF
50-59 years, male
951.5
GTEX-XMK1-2026-SM-4B65K
40-49 years, male
951.5
GTEX-U4B1-1526-SM-4DXSL
40-49 years, male
950.5
GTEX-11NSD-1026-SM-5N9BE
20-29 years, male
946.7
GTEX-XBEC-0126-SM-4GIDT
50-59 years, male
943.6
GTEX-14BIL-2226-SM-73KWF
50-59 years, male
943.5
GTEX-14C39-0526-SM-664OF
40-49 years, male
943.5
GTEX-131XE-0426-SM-5IJF4
50-59 years, male
941.9
GTEX-PLZ5-1526-SM-3P5ZX
50-59 years, male
941.1
GTEX-13O61-2026-SM-5J2M6
60-69 years, male
941.0
GTEX-1AX9J-1226-SM-72D6P
60-69 years, male
940.2
GTEX-145MF-1726-SM-5LU9H
50-59 years, male
938.0
GTEX-12WSI-2126-SM-5GCMV
50-59 years, male
937.5
GTEX-14A5H-0626-SM-5TDCO
60-69 years, male
934.5
GTEX-X261-2326-SM-4PQYU
50-59 years, male
933.8
GTEX-12ZZY-0126-SM-5LZV2
60-69 years, male
931.7
GTEX-O5YT-2126-SM-3MJGD
20-29 years, male
929.9
GTEX-RN64-2326-SM-48FDW
50-59 years, male
925.9
GTEX-RUSQ-2126-SM-47JXK
50-59 years, male
923.7
GTEX-Y5V6-1726-SM-4VDSZ
60-69 years, male
923.7
GTEX-1AX9I-1826-SM-72D5I
60-69 years, male
921.8
GTEX-WFON-2026-SM-4LVMW
40-49 years, male
921.3
GTEX-117YX-2026-SM-5GIEF
50-59 years, male
918.4
GTEX-N7MS-0126-SM-3TW8O
60-69 years, male
917.1
GTEX-QEG4-0126-SM-48TZE
30-39 years, male
915.8
GTEX-1CB4G-2426-SM-7DUGM
60-69 years, male
915.3
GTEX-13OVH-0726-SM-5N9BU
70-79 years, male
915.2
GTEX-132QS-1226-SM-5P9GD
60-69 years, male
914.9
GTEX-WK11-0326-SM-4OOS6
50-59 years, male
914.0
GTEX-13FHP-2826-SM-5IJFW
50-59 years, male
909.5
GTEX-14753-0626-SM-5Q5CY
60-69 years, male
909.3
GTEX-1C6VR-1526-SM-7MKG9
60-69 years, male
908.5
GTEX-16Z82-2726-SM-7KULV
60-69 years, male
902.9
GTEX-ZAB5-2426-SM-5CVMW
50-59 years, male
902.2
GTEX-1212Z-0326-SM-5FQSJ
60-69 years, male
901.0
GTEX-ZVTK-0126-SM-57WDG
20-29 years, male
900.0
GTEX-OOBK-2126-SM-3LK5T
40-49 years, male
897.1
GTEX-13O1R-0726-SM-5IJEI
60-69 years, male
894.4
GTEX-14DAR-1126-SM-793AT
50-59 years, male
891.3
GTEX-13OVK-2226-SM-6LPJY
40-49 years, male
891.1
GTEX-Q2AI-1226-SM-48U14
40-49 years, male
889.4
GTEX-11WQC-2326-SM-5EQKE
60-69 years, male
888.7
GTEX-13VXU-0726-SM-5J2O7
50-59 years, male
884.3
GTEX-ZPU1-2126-SM-57WED
40-49 years, male
883.7
GTEX-147JS-0126-SM-5S2TW
60-69 years, male
882.7
GTEX-1CB4I-0926-SM-7MKFY
50-59 years, male
881.7
GTEX-S7PM-0626-SM-4AD4Q
60-69 years, male
880.9
GTEX-14E1K-1926-SM-73KWS
50-59 years, male
878.6
GTEX-111CU-1726-SM-5EGHM
50-59 years, male
878.5
GTEX-OIZI-0126-SM-3NB13
40-49 years, male
877.0
GTEX-12WSH-0326-SM-5GCNH
50-59 years, male
874.2
GTEX-1E1VI-1026-SM-7MKGQ
50-59 years, male
871.5
GTEX-12BJ1-1326-SM-5BC5P
60-69 years, male
870.5
GTEX-18A66-0726-SM-72D77
60-69 years, male
870.5
GTEX-U8XE-0126-SM-4E3I3
30-39 years, male
869.5
GTEX-144GM-0426-SM-5Q5C8
20-29 years, male
868.1
GTEX-13FLW-2126-SM-5N9FD
50-59 years, male
867.8
GTEX-VJYA-1426-SM-4KL1Y
60-69 years, male
866.9
GTEX-11NV4-1726-SM-5N9FC
60-69 years, male
865.7
GTEX-12696-0226-SM-5EGL3
60-69 years, male
862.8
GTEX-RWSA-2426-SM-47JXR
40-49 years, male
862.8
GTEX-15RIF-1826-SM-6M469
20-29 years, male
862.5
GTEX-12WSL-2326-SM-5DUXQ
50-59 years, male
858.5
GTEX-11P7K-1026-SM-5HL6Y
30-39 years, male
857.7
GTEX-S4Q7-1226-SM-4AD5I
20-29 years, male
857.2
GTEX-Y8E4-2226-SM-5LU94
40-49 years, male
853.7
GTEX-WZTO-0326-SM-4PQYZ
40-49 years, male
853.2
GTEX-139T6-1226-SM-5IFFC
50-59 years, male
851.0
GTEX-11OF3-1826-SM-5987N
60-69 years, male
850.9
GTEX-ZVZP-2226-SM-57WBF
50-59 years, male
850.7
GTEX-QLQ7-1426-SM-4R1JX
60-69 years, male
850.3
GTEX-13OW5-2526-SM-5L3I1
60-69 years, male
849.7
GTEX-OXRL-2126-SM-3NM98
50-59 years, male
849.3
GTEX-WH7G-1926-SM-4LVMM
40-49 years, male
849.1
GTEX-14PJ2-2026-SM-6AJAQ
50-59 years, male
848.4
GTEX-11DXZ-2126-SM-59881
50-59 years, male
845.7
GTEX-ZTSS-1526-SM-51MTC
40-49 years, male
842.5
GTEX-13NZA-2526-SM-5IJFX
60-69 years, male
841.8
GTEX-16XZZ-0626-SM-6M47T
60-69 years, male
827.8
GTEX-13NZB-2026-SM-5MR4M
30-39 years, male
825.1
GTEX-X5EB-2026-SM-4E3KA
40-49 years, male
823.7
GTEX-1A3MV-0726-SM-72D5M
30-39 years, male
823.0
GTEX-14B4R-1026-SM-5TDDS
30-39 years, male
820.3
GTEX-13O21-1226-SM-5J2MK
50-59 years, male
819.9
GTEX-OHPM-2126-SM-3LK75
50-59 years, male
819.5
GTEX-111VG-1926-SM-5GIDO
60-69 years, male
818.4
GTEX-1EKGG-2726-SM-79OOD
60-69 years, male
818.2
GTEX-13OVL-0426-SM-5IFG6
50-59 years, male
816.8
GTEX-13111-1526-SM-5EGJX
50-59 years, male
816.7
GTEX-ZUA1-2726-SM-59HLJ
40-49 years, male
816.1
GTEX-14PJN-1626-SM-68727
30-39 years, male
809.1
GTEX-11EQ9-1926-SM-5PNVV
30-39 years, male
807.8
GTEX-145MH-2326-SM-5O9AW
50-59 years, male
806.8
GTEX-ZT9W-2226-SM-57WFU
50-59 years, male
805.6
GTEX-V1D1-2126-SM-4JBH4
50-59 years, male
803.2
GTEX-11NUK-2626-SM-5A5MB
50-59 years, male
802.5
GTEX-ZDYS-1326-SM-5IJFF
50-59 years, male
800.8
GTEX-14E7W-0726-SM-664OK
70-79 years, male
796.5
GTEX-Z93S-1726-SM-5HL8G
50-59 years, male
796.3
GTEX-12C56-1426-SM-5FQSW
20-29 years, male
794.1
GTEX-ZTX8-1126-SM-51MRM
20-29 years, male
794.1
GTEX-Y111-2426-SM-4TT23
50-59 years, male
790.9
GTEX-YB5E-1926-SM-5IFIG
40-49 years, male
788.3
GTEX-XAJ8-1326-SM-47JYT
40-49 years, male
787.7
GTEX-ZZ64-1126-SM-5GZXY
20-29 years, male
783.2
GTEX-16MTA-1126-SM-6LPJA
50-59 years, male
781.7
GTEX-111YS-2026-SM-5EGGL
60-69 years, male
780.6
GTEX-13FTW-1326-SM-5LZZD
40-49 years, male
774.4
GTEX-17F9E-1626-SM-7EWDB
50-59 years, male
773.4
GTEX-117XS-2026-SM-5GID1
60-69 years, male
772.0
GTEX-14PJ3-1626-SM-664O5
50-59 years, male
769.4
GTEX-1CAMR-1526-SM-79OLP
20-29 years, male
767.8
GTEX-14PHX-1426-SM-69LPP
50-59 years, male
764.0
GTEX-1EU9M-1626-SM-7EWF2
60-69 years, male
760.0
GTEX-18QFQ-1926-SM-72D5H
30-39 years, male
755.4
GTEX-11TUW-2226-SM-5EQL9
60-69 years, male
743.1
GTEX-14C5O-2326-SM-73KYU
60-69 years, male
739.2
GTEX-17MFQ-0826-SM-793C8
20-29 years, male
736.9
GTEX-1B8KE-1426-SM-7EWEL
50-59 years, male
734.5
GTEX-11LCK-2326-SM-5HL53
30-39 years, male
721.4
GTEX-ZYT6-2726-SM-5GICP
30-39 years, male
711.7
GTEX-117YW-1526-SM-5EGGP
50-59 years, male
707.7
GTEX-11P82-1526-SM-5BC5M
20-29 years, male
695.6
GTEX-11ZUS-2726-SM-5FQUA
30-39 years, male
692.8
GTEX-1C64N-2826-SM-7IGP5
50-59 years, male
688.6
GTEX-YEC3-1726-SM-5IFIK
50-59 years, male
661.0
GTEX-16AAH-1626-SM-7EWE2
50-59 years, male
650.5
GTEX-111FC-1926-SM-5GZYC
60-69 years, male
642.3
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TESTIS - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.