We use cookies to enhance the usability of our website. If you continue, we'll assume that you are happy to receive all cookies. More information. Don't show this again.
On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression (NX) levels for human tissues was created by combining the data from these three transcriptomics datasets (HPA, GTEx and FANTOM5).
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Antibody staining in the annotated cell types in the current human tissue is reported as not detected, low, medium, or high. This score is based on the staining intensity and fraction of stained cells.
The RNA-seq details section shows detailed information about the individual samples used for the transcript profiling and results of the RNA-seq analysis.
Information about each individual sample is listed below, including gender, age, a tissue section image and estimated fractions of cell types. pTPM (transcripts per million) values give a quantification of the gene abundance which is comparable between different genes and samples.
Cells in seminiferous ducts: 65 Other cell types: 35
Male, age 34
Testis sample 278 pTPM: 566
Cell types%
Cells in seminiferous ducts: 75 Other cell types: 25
Male, age 62
Testis sample 279 pTPM: 612.1
Cell types%
Cells in seminiferous ducts: 70 Other cell types: 30
Male, age 34
Testis sample 280 pTPM: 1045.7
Cell types%
Cells in seminiferous ducts: 60 Other cell types: 40
Male, age 26
Testis sample 281 pTPM: 507.4
Cell types%
Cells in seminiferous ducts: 55 Other cell types: 45
Male, age 50
Testis sample 282 pTPM: 460.4
Cell types%
Cells in seminiferous ducts: 85 Other cell types: 15
Male, age 37
Testis sample 283 pTPM: 618.7
Cell types%
Cells in seminiferous ducts: 35 Other cell types: 65
Male, age 51
Testis sample 349 pTPM: 819.8
Cell types%
Cells in seminiferous ducts: 65 Other cell types: 35
Male, age 34
Testis sample 354 pTPM: 822.6
Cell types%
Cells in seminiferous ducts: 70 Other cell types: 30
Male, age 26
Testis sample 371 pTPM: 1006.3
Cell types%
Cells in seminiferous ducts: 75 Other cell types: 25
TESTIS - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as average pTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. pTPM values of the individual samples are presented next to the box plot.
Max pTPM: 3135.5
Min pTPM: 290.2
Std pTPM: 282.8
Median pTPM: 571.8
GTEX-NPJ8-1226-SM-3MJHM
40-49 years, male
3135.5
GTEX-1C6VQ-1426-SM-73KTW
50-59 years, male
1889.7
GTEX-148VJ-2526-SM-5TDEI
70-79 years, male
1745.5
GTEX-PLZ6-1226-SM-3P5ZS
30-39 years, male
1539.1
GTEX-1399T-1526-SM-5P9J6
60-69 years, male
1527.2
GTEX-17F97-2726-SM-7IGQ4
50-59 years, male
1467.5
GTEX-18A67-1826-SM-7KFT7
50-59 years, male
1413.8
GTEX-1C6VS-1126-SM-7EWEW
60-69 years, male
1300.8
GTEX-12WSM-1326-SM-5GCP9
60-69 years, male
1297.5
GTEX-Y3I4-2026-SM-4TT6Z
50-59 years, male
1289.5
GTEX-1B8KZ-1826-SM-73KVJ
50-59 years, male
1233.0
GTEX-13QJ3-0226-SM-5S2PU
50-59 years, male
1200.8
GTEX-1E2YA-2826-SM-7EPIN
50-59 years, male
1164.1
GTEX-1CB4J-1626-SM-79OOK
60-69 years, male
1156.0
GTEX-1AX8Z-2826-SM-73KTZ
60-69 years, male
1147.6
GTEX-11DXY-0226-SM-5H123
60-69 years, male
1143.3
GTEX-ZLFU-2026-SM-4WWG2
40-49 years, male
1093.0
GTEX-145LT-0426-SM-5LUAP
40-49 years, male
1090.8
GTEX-147GR-0626-SM-5S2PK
60-69 years, male
1076.8
GTEX-S4Z8-2126-SM-4AD5H
60-69 years, male
1051.4
GTEX-13OW8-0526-SM-5KM24
60-69 years, male
1047.7
GTEX-WVJS-2726-SM-4MVNU
50-59 years, male
1021.3
GTEX-1399R-1626-SM-5P9GG
30-39 years, male
970.6
GTEX-145MF-1726-SM-5LU9H
50-59 years, male
958.7
GTEX-147F4-0626-SM-5LUAK
50-59 years, male
937.6
GTEX-YJ89-0626-SM-4TT3Z
60-69 years, male
925.0
GTEX-14A6H-2326-SM-5Q5B5
30-39 years, male
917.0
GTEX-WY7C-2226-SM-4ONCS
50-59 years, male
899.6
GTEX-ZAB4-0126-SM-5CVMG
40-49 years, male
896.7
GTEX-13OW6-0126-SM-5IJGM
50-59 years, male
890.9
GTEX-X3Y1-2626-SM-4PQZI
40-49 years, male
879.7
GTEX-NFK9-0126-SM-3LK5H
40-49 years, male
846.4
GTEX-QEG5-0126-SM-4R1JR
20-29 years, male
820.6
GTEX-11ONC-2226-SM-5HL6D
60-69 years, male
818.2
GTEX-1BAJH-0726-SM-7IGMF
70-79 years, male
811.9
GTEX-117YX-2026-SM-5GIEF
50-59 years, male
777.6
GTEX-S3XE-1526-SM-4AD5A
50-59 years, male
773.8
GTEX-U8T8-1126-SM-4DXUE
60-69 years, male
755.3
GTEX-PW2O-1426-SM-48TCD
20-29 years, male
754.2
GTEX-13NYB-2226-SM-5MR58
40-49 years, male
753.5
GTEX-1AYCT-1026-SM-79ONR
50-59 years, male
753.2
GTEX-17HGU-0726-SM-7DUFN
50-59 years, male
751.8
GTEX-WFG8-1926-SM-4LVM1
20-29 years, male
736.6
GTEX-OIZI-0126-SM-3NB13
40-49 years, male
731.0
GTEX-14A5H-0626-SM-5TDCO
60-69 years, male
730.9
GTEX-18A7B-1526-SM-7KFTH
70-79 years, male
726.0
GTEX-15RJ7-1826-SM-7KUMJ
40-49 years, male
725.7
GTEX-14C38-0126-SM-5YY9V
50-59 years, male
725.4
GTEX-S33H-0126-SM-4AD62
30-39 years, male
724.8
GTEX-ZDTT-2026-SM-5K7TY
60-69 years, male
719.0
GTEX-12BJ1-1326-SM-5BC5P
60-69 years, male
714.9
GTEX-Y9LG-1726-SM-4VBQE
30-39 years, male
713.2
GTEX-11TT1-2226-SM-5GU6B
20-29 years, male
712.4
GTEX-13N2G-0126-SM-5N9DV
60-69 years, male
710.4
GTEX-VJYA-1426-SM-4KL1Y
60-69 years, male
697.3
GTEX-S7SE-0326-SM-4AT5Q
50-59 years, male
691.0
GTEX-RN64-2326-SM-48FDW
50-59 years, male
689.2
GTEX-YF7O-2026-SM-4W1YE
50-59 years, male
686.9
GTEX-14B4R-1026-SM-5TDDS
30-39 years, male
685.3
GTEX-11P7K-1026-SM-5HL6Y
30-39 years, male
685.1
GTEX-QV44-1726-SM-4R1KG
50-59 years, male
681.7
GTEX-OIZH-2126-SM-3NB1P
50-59 years, male
681.2
GTEX-V955-1926-SM-4KL1L
60-69 years, male
680.0
GTEX-15SKB-0326-SM-6M477
60-69 years, male
675.0
GTEX-Y8E4-2226-SM-5LU94
40-49 years, male
671.5
GTEX-ZQUD-2026-SM-51MSM
30-39 years, male
670.5
GTEX-14PJ4-1726-SM-664OC
60-69 years, male
669.1
GTEX-XBED-2026-SM-4AT5D
60-69 years, male
668.9
GTEX-XPT6-1626-SM-4B655
20-29 years, male
666.3
GTEX-T5JC-0726-SM-4DM55
20-29 years, male
665.5
GTEX-ZYFC-0126-SM-5GIEH
50-59 years, male
660.8
GTEX-139TT-2226-SM-5LZWO
60-69 years, male
660.0
GTEX-YB5E-1926-SM-5IFIG
40-49 years, male
658.6
GTEX-Q2AH-1526-SM-48TZG
40-49 years, male
658.2
GTEX-OOBJ-2126-SM-3NB1N
60-69 years, male
657.8
GTEX-XGQ4-2026-SM-4AT6G
50-59 years, male
655.0
GTEX-14E7W-0726-SM-664OK
70-79 years, male
653.2
GTEX-1A3MX-1026-SM-731F8
60-69 years, male
651.9
GTEX-144GN-1626-SM-5Q5BU
50-59 years, male
647.3
GTEX-P4QS-2126-SM-3NMCF
60-69 years, male
645.9
GTEX-ZT9X-1426-SM-5DUX1
40-49 years, male
645.2
GTEX-1AX9K-1826-SM-731CY
50-59 years, male
645.1
GTEX-QLQ7-1426-SM-4R1JX
60-69 years, male
645.0
GTEX-145MO-0126-SM-5S2QU
60-69 years, male
643.1
GTEX-ZVTK-0126-SM-57WDG
20-29 years, male
642.4
GTEX-X261-2326-SM-4PQYU
50-59 years, male
642.2
GTEX-11EI6-2226-SM-5EGJM
60-69 years, male
641.9
GTEX-14PHX-1426-SM-69LPP
50-59 years, male
637.7
GTEX-U3ZH-1526-SM-4DXV1
30-39 years, male
628.3
GTEX-1339X-1926-SM-5PNVP
40-49 years, male
625.6
GTEX-17KNJ-1126-SM-7KFT6
50-59 years, male
623.3
GTEX-ZV7C-2026-SM-5NQ8F
50-59 years, male
622.2
GTEX-O5YT-2126-SM-3MJGD
20-29 years, male
621.8
GTEX-R55C-1426-SM-48FED
40-49 years, male
620.9
GTEX-1F6IF-0526-SM-7MKHD
60-69 years, male
616.3
GTEX-XMK1-2026-SM-4B65K
40-49 years, male
611.1
GTEX-17F9E-1626-SM-7EWDB
50-59 years, male
610.0
GTEX-YEC4-1526-SM-4W1YU
40-49 years, male
608.0
GTEX-PLZ5-1526-SM-3P5ZX
50-59 years, male
606.5
GTEX-QMRM-1526-SM-4R1K6
50-59 years, male
605.2
GTEX-QLQW-1026-SM-447A9
30-39 years, male
602.0
GTEX-131XE-0426-SM-5IJF4
50-59 years, male
600.6
GTEX-XPVG-2226-SM-4B65U
50-59 years, male
596.2
GTEX-U4B1-1526-SM-4DXSL
40-49 years, male
595.9
GTEX-WVLH-2626-SM-4MVNV
50-59 years, male
594.7
GTEX-TKQ1-0926-SM-4DXU2
30-39 years, male
593.8
GTEX-12WSI-2126-SM-5GCMV
50-59 years, male
593.4
GTEX-139TS-1726-SM-5IJG5
60-69 years, male
593.4
GTEX-14E6E-1026-SM-664N9
30-39 years, male
593.0
GTEX-R55D-0126-SM-48FEL
50-59 years, male
592.5
GTEX-13OVH-0726-SM-5N9BU
70-79 years, male
590.4
GTEX-RWSA-2426-SM-47JXR
40-49 years, male
588.9
GTEX-RUSQ-2126-SM-47JXK
50-59 years, male
587.9
GTEX-16YQH-1826-SM-6LPJW
30-39 years, male
587.0
GTEX-183FY-1926-SM-7KFRI
20-29 years, male
586.6
GTEX-11GS4-2026-SM-5N9CP
60-69 years, male
586.1
GTEX-144GL-0726-SM-5LU4P
50-59 years, male
582.6
GTEX-16Z82-2726-SM-7KULV
60-69 years, male
582.4
GTEX-WHSB-2126-SM-4M1XF
50-59 years, male
581.9
GTEX-1C64O-2126-SM-7IGPB
60-69 years, male
581.5
GTEX-14753-0626-SM-5Q5CY
60-69 years, male
581.3
GTEX-13NZB-2026-SM-5MR4M
30-39 years, male
580.8
GTEX-16XZZ-0626-SM-6M47T
60-69 years, male
580.4
GTEX-S95S-1126-SM-4B64E
60-69 years, male
580.1
GTEX-1B996-2326-SM-731EC
60-69 years, male
578.9
GTEX-1CB4I-0926-SM-7MKFY
50-59 years, male
577.3
GTEX-WFON-2026-SM-4LVMW
40-49 years, male
574.6
GTEX-T8EM-0126-SM-4DM5R
40-49 years, male
574.0
GTEX-RM2N-1326-SM-48FCW
50-59 years, male
573.2
GTEX-U3ZM-1626-SM-4DXSK
40-49 years, male
571.8
GTEX-11O72-0726-SM-5P9GO
40-49 years, male
569.7
GTEX-ZVZP-2226-SM-57WBF
50-59 years, male
568.5
GTEX-147JS-0126-SM-5S2TW
60-69 years, male
565.4
GTEX-18D9B-2726-SM-72D73
60-69 years, male
564.8
GTEX-R55E-0726-SM-48FCZ
20-29 years, male
563.2
GTEX-12WSL-2326-SM-5DUXQ
50-59 years, male
562.5
GTEX-Z93S-1726-SM-5HL8G
50-59 years, male
561.6
GTEX-13FTW-1326-SM-5LZZD
40-49 years, male
559.4
GTEX-13N1W-2626-SM-5IJEP
70-79 years, male
558.6
GTEX-11NSD-1026-SM-5N9BE
20-29 years, male
558.4
GTEX-14DAR-1126-SM-793AT
50-59 years, male
558.3
GTEX-Y5V6-1726-SM-4VDSZ
60-69 years, male
557.7
GTEX-XLM4-1526-SM-4AT6D
60-69 years, male
557.3
GTEX-139T6-1226-SM-5IFFC
50-59 years, male
556.4
GTEX-ZTX8-1126-SM-51MRM
20-29 years, male
556.3
GTEX-WHSE-0426-SM-4M1XO
20-29 years, male
554.9
GTEX-13O61-2026-SM-5J2M6
60-69 years, male
554.5
GTEX-18A6Q-2626-SM-7KFT3
60-69 years, male
549.9
GTEX-YFCO-1726-SM-4W21S
40-49 years, male
549.2
GTEX-15SHV-1926-SM-6LPI1
60-69 years, male
549.0
GTEX-1E1VI-1026-SM-7MKGQ
50-59 years, male
547.9
GTEX-WYJK-1826-SM-4ONDM
50-59 years, male
547.5
GTEX-14PJN-1626-SM-68727
30-39 years, male
544.7
GTEX-ZAB5-2426-SM-5CVMW
50-59 years, male
543.3
GTEX-ZZ64-1126-SM-5GZXY
20-29 years, male
542.9
GTEX-15RIE-0626-SM-6M47G
60-69 years, male
542.5
GTEX-14C39-0526-SM-664OF
40-49 years, male
542.2
GTEX-ZA64-1626-SM-5CVME
20-29 years, male
540.7
GTEX-12ZZY-0126-SM-5LZV2
60-69 years, male
539.6
GTEX-14BIL-2226-SM-73KWF
50-59 years, male
538.1
GTEX-N7MS-0126-SM-3TW8O
60-69 years, male
538.1
GTEX-REY6-0126-SM-48FDT
60-69 years, male
533.5
GTEX-1212Z-0326-SM-5FQSJ
60-69 years, male
533.2
GTEX-ZUA1-2726-SM-59HLJ
40-49 years, male
528.2
GTEX-117XS-2026-SM-5GID1
60-69 years, male
527.9
GTEX-18465-1526-SM-7KFTV
50-59 years, male
524.3
GTEX-145MH-2326-SM-5O9AW
50-59 years, male
524.1
GTEX-17HHY-0726-SM-7IGM6
50-59 years, male
521.4
GTEX-XBEC-0126-SM-4GIDT
50-59 years, male
520.8
GTEX-132QS-1226-SM-5P9GD
60-69 years, male
520.3
GTEX-UPJH-0126-SM-4IHLL
50-59 years, male
519.0
GTEX-16MTA-1126-SM-6LPJA
50-59 years, male
518.7
GTEX-13O1R-0726-SM-5IJEI
60-69 years, male
517.2
GTEX-17MF6-2726-SM-7IGMZ
40-49 years, male
515.5
GTEX-1GMRU-0626-SM-7MKH2
50-59 years, male
515.3
GTEX-QV31-1126-SM-4R1K4
30-39 years, male
515.2
GTEX-WH7G-1926-SM-4LVMM
40-49 years, male
515.2
GTEX-17GQL-1726-SM-718B8
60-69 years, male
513.4
GTEX-111CU-1726-SM-5EGHM
50-59 years, male
511.0
GTEX-15RIF-1826-SM-6M469
20-29 years, male
511.0
GTEX-13NZA-2526-SM-5IJFX
60-69 years, male
510.7
GTEX-SNMC-1026-SM-4DM7K
20-29 years, male
510.1
GTEX-14PJ3-1626-SM-664O5
50-59 years, male
507.7
GTEX-1AX9I-1826-SM-72D5I
60-69 years, male
507.6
GTEX-13FLW-2126-SM-5N9FD
50-59 years, male
507.0
GTEX-12WSH-0326-SM-5GCNH
50-59 years, male
506.5
GTEX-TKQ2-1526-SM-4DXUN
50-59 years, male
506.2
GTEX-S4Q7-1226-SM-4AD5I
20-29 years, male
504.6
GTEX-U8XE-0126-SM-4E3I3
30-39 years, male
503.3
GTEX-11P82-1526-SM-5BC5M
20-29 years, male
503.0
GTEX-V1D1-2126-SM-4JBH4
50-59 years, male
502.4
GTEX-SNOS-1126-SM-4DM67
40-49 years, male
500.2
GTEX-13VXU-0726-SM-5J2O7
50-59 years, male
498.0
GTEX-12696-0226-SM-5EGL3
60-69 years, male
496.1
GTEX-ZYT6-2726-SM-5GICP
30-39 years, male
494.0
GTEX-13OVK-2226-SM-6LPJY
40-49 years, male
493.3
GTEX-15G19-1326-SM-6LPIR
60-69 years, male
492.9
GTEX-11DXZ-2126-SM-59881
50-59 years, male
490.6
GTEX-14ABY-0626-SM-5Q5C9
50-59 years, male
488.6
GTEX-1B8SF-2926-SM-731DL
60-69 years, male
487.8
GTEX-11NUK-2626-SM-5A5MB
50-59 years, male
486.3
GTEX-1AX9J-1226-SM-72D6P
60-69 years, male
485.9
GTEX-13OW5-2526-SM-5L3I1
60-69 years, male
485.7
GTEX-11OF3-1826-SM-5987N
60-69 years, male
484.5
GTEX-ZDYS-1326-SM-5IJFF
50-59 years, male
483.3
GTEX-17HHE-1226-SM-793C6
50-59 years, male
482.8
GTEX-12C56-1426-SM-5FQSW
20-29 years, male
482.4
GTEX-18QFQ-1926-SM-72D5H
30-39 years, male
480.1
GTEX-ZPU1-2126-SM-57WED
40-49 years, male
479.1
GTEX-ZT9W-2226-SM-57WFU
50-59 years, male
476.3
GTEX-14E1K-1926-SM-73KWS
50-59 years, male
472.5
GTEX-18A66-0726-SM-72D77
60-69 years, male
471.6
GTEX-1CAMR-1526-SM-79OLP
20-29 years, male
470.2
GTEX-XQ3S-2726-SM-4BOP2
20-29 years, male
468.8
GTEX-1B8KE-1426-SM-7EWEL
50-59 years, male
468.2
GTEX-13111-1526-SM-5EGJX
50-59 years, male
467.7
GTEX-11EQ8-1426-SM-5EGJR
60-69 years, male
466.5
GTEX-13O21-1226-SM-5J2MK
50-59 years, male
464.8
GTEX-17HII-0126-SM-7KFSS
50-59 years, male
461.2
GTEX-11EQ9-1926-SM-5PNVV
30-39 years, male
459.5
GTEX-T6MN-2026-SM-4DM7L
50-59 years, male
459.5
GTEX-Q2AI-1226-SM-48U14
40-49 years, male
456.8
GTEX-OXRL-2126-SM-3NM98
50-59 years, male
452.2
GTEX-144GM-0426-SM-5Q5C8
20-29 years, male
451.7
GTEX-13OVL-0426-SM-5IFG6
50-59 years, male
450.0
GTEX-WOFM-1126-SM-4OOSB
20-29 years, male
449.7
GTEX-S7PM-0626-SM-4AD4Q
60-69 years, male
440.5
GTEX-XAJ8-1326-SM-47JYT
40-49 years, male
430.2
GTEX-1C4CL-1926-SM-731DY
30-39 years, male
428.4
GTEX-X5EB-2026-SM-4E3KA
40-49 years, male
425.4
GTEX-111YS-2026-SM-5EGGL
60-69 years, male
425.0
GTEX-117YW-1526-SM-5EGGP
50-59 years, male
422.5
GTEX-13112-0226-SM-5P9IV
50-59 years, male
422.1
GTEX-14PJ2-2026-SM-6AJAQ
50-59 years, male
421.1
GTEX-111FC-1926-SM-5GZYC
60-69 years, male
419.4
GTEX-ZTSS-1526-SM-51MTC
40-49 years, male
419.4
GTEX-1EU9M-1626-SM-7EWF2
60-69 years, male
419.2
GTEX-11WQC-2326-SM-5EQKE
60-69 years, male
415.5
GTEX-11NV4-1726-SM-5N9FC
60-69 years, male
414.5
GTEX-OOBK-2126-SM-3LK5T
40-49 years, male
412.2
GTEX-11LCK-2326-SM-5HL53
30-39 years, male
411.6
GTEX-1CB4G-2426-SM-7DUGM
60-69 years, male
408.3
GTEX-1A3MV-0726-SM-72D5M
30-39 years, male
408.1
GTEX-WZTO-0326-SM-4PQYZ
40-49 years, male
406.9
GTEX-Y111-2426-SM-4TT23
50-59 years, male
406.3
GTEX-16AAH-1626-SM-7EWE2
50-59 years, male
404.8
GTEX-14C5O-2326-SM-73KYU
60-69 years, male
404.6
GTEX-YEC3-1726-SM-5IFIK
50-59 years, male
401.9
GTEX-QEG4-0126-SM-48TZE
30-39 years, male
401.8
GTEX-WK11-0326-SM-4OOS6
50-59 years, male
401.6
GTEX-1C6VR-1526-SM-7MKG9
60-69 years, male
398.0
GTEX-OHPM-2126-SM-3LK75
50-59 years, male
395.3
GTEX-11TUW-2226-SM-5EQL9
60-69 years, male
389.8
GTEX-17MFQ-0826-SM-793C8
20-29 years, male
388.2
GTEX-1EKGG-2726-SM-79OOD
60-69 years, male
386.3
GTEX-111VG-1926-SM-5GIDO
60-69 years, male
367.3
GTEX-1C64N-2826-SM-7IGP5
50-59 years, male
316.5
GTEX-13FHP-2826-SM-5IJFW
50-59 years, male
315.8
GTEX-11ZUS-2726-SM-5FQUA
30-39 years, male
290.2
Show allShow less
TESTIS - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.