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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression (NX) levels for human tissues was created by combining the data from these three transcriptomics datasets (HPA, GTEx and FANTOM5).
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Antibody staining in the annotated cell types in the current human tissue is reported as not detected, low, medium, or high. This score is based on the staining intensity and fraction of stained cells.
The RNA-seq details section shows detailed information about the individual samples used for the transcript profiling and results of the RNA-seq analysis.
Information about each individual sample is listed below, including gender, age, a tissue section image and estimated fractions of cell types. pTPM (transcripts per million) values give a quantification of the gene abundance which is comparable between different genes and samples.
Cells in seminiferous ducts: 65 Other cell types: 35
Male, age 34
Testis sample 278 pTPM: 963.8
Cell types%
Cells in seminiferous ducts: 75 Other cell types: 25
Male, age 62
Testis sample 279 pTPM: 941.1
Cell types%
Cells in seminiferous ducts: 70 Other cell types: 30
Male, age 34
Testis sample 280 pTPM: 973.3
Cell types%
Cells in seminiferous ducts: 60 Other cell types: 40
Male, age 26
Testis sample 281 pTPM: 768.2
Cell types%
Cells in seminiferous ducts: 55 Other cell types: 45
Male, age 50
Testis sample 282 pTPM: 841.9
Cell types%
Cells in seminiferous ducts: 85 Other cell types: 15
Male, age 37
Testis sample 283 pTPM: 825.6
Cell types%
Cells in seminiferous ducts: 35 Other cell types: 65
Male, age 51
Testis sample 349 pTPM: 158.3
Cell types%
Cells in seminiferous ducts: 65 Other cell types: 35
Male, age 34
Testis sample 354 pTPM: 472.9
Cell types%
Cells in seminiferous ducts: 70 Other cell types: 30
Male, age 26
Testis sample 371 pTPM: 290.2
Cell types%
Cells in seminiferous ducts: 75 Other cell types: 25
TESTIS - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as average pTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. pTPM values of the individual samples are presented next to the box plot.
Max pTPM: 1241.7
Min pTPM: 555.8
Std pTPM: 121.4
Median pTPM: 945.8
GTEX-UPJH-0126-SM-4IHLL
50-59 years, male
1241.7
GTEX-18465-1526-SM-7KFTV
50-59 years, male
1238.5
GTEX-XBED-2026-SM-4AT5D
60-69 years, male
1202.1
GTEX-YF7O-2026-SM-4W1YE
50-59 years, male
1180.6
GTEX-RM2N-1326-SM-48FCW
50-59 years, male
1179.0
GTEX-PLZ5-1526-SM-3P5ZX
50-59 years, male
1175.8
GTEX-13O21-1226-SM-5J2MK
50-59 years, male
1172.1
GTEX-S3XE-1526-SM-4AD5A
50-59 years, male
1147.7
GTEX-16YQH-1826-SM-6LPJW
30-39 years, male
1144.3
GTEX-13O61-2026-SM-5J2M6
60-69 years, male
1139.9
GTEX-ZDTT-2026-SM-5K7TY
60-69 years, male
1136.1
GTEX-XQ3S-2726-SM-4BOP2
20-29 years, male
1135.1
GTEX-QLQW-1026-SM-447A9
30-39 years, male
1133.4
GTEX-WFG8-1926-SM-4LVM1
20-29 years, male
1130.2
GTEX-WVLH-2626-SM-4MVNV
50-59 years, male
1128.8
GTEX-14E6E-1026-SM-664N9
30-39 years, male
1128.6
GTEX-YFCO-1726-SM-4W21S
40-49 years, male
1125.2
GTEX-T8EM-0126-SM-4DM5R
40-49 years, male
1117.8
GTEX-1AYCT-1026-SM-79ONR
50-59 years, male
1115.7
GTEX-WY7C-2226-SM-4ONCS
50-59 years, male
1115.7
GTEX-117YX-2026-SM-5GIEF
50-59 years, male
1112.4
GTEX-11EI6-2226-SM-5EGJM
60-69 years, male
1111.6
GTEX-1AX9J-1226-SM-72D6P
60-69 years, male
1109.5
GTEX-S7SE-0326-SM-4AT5Q
50-59 years, male
1106.5
GTEX-1AX9K-1826-SM-731CY
50-59 years, male
1104.0
GTEX-14C38-0126-SM-5YY9V
50-59 years, male
1102.6
GTEX-SNMC-1026-SM-4DM7K
20-29 years, male
1099.8
GTEX-WFON-2026-SM-4LVMW
40-49 years, male
1094.5
GTEX-13FTW-1326-SM-5LZZD
40-49 years, male
1091.6
GTEX-WK11-0326-SM-4OOS6
50-59 years, male
1080.8
GTEX-S33H-0126-SM-4AD62
30-39 years, male
1080.3
GTEX-OHPM-2126-SM-3LK75
50-59 years, male
1079.0
GTEX-U3ZH-1526-SM-4DXV1
30-39 years, male
1078.6
GTEX-TKQ1-0926-SM-4DXU2
30-39 years, male
1078.5
GTEX-O5YT-2126-SM-3MJGD
20-29 years, male
1072.4
GTEX-18A66-0726-SM-72D77
60-69 years, male
1071.5
GTEX-X261-2326-SM-4PQYU
50-59 years, male
1071.1
GTEX-13NZB-2026-SM-5MR4M
30-39 years, male
1068.9
GTEX-ZTSS-1526-SM-51MTC
40-49 years, male
1065.0
GTEX-13VXU-0726-SM-5J2O7
50-59 years, male
1064.0
GTEX-OOBK-2126-SM-3LK5T
40-49 years, male
1063.3
GTEX-14BIL-2226-SM-73KWF
50-59 years, male
1062.9
GTEX-17KNJ-1126-SM-7KFT6
50-59 years, male
1061.5
GTEX-139TS-1726-SM-5IJG5
60-69 years, male
1058.0
GTEX-13NZA-2526-SM-5IJFX
60-69 years, male
1057.8
GTEX-1CB4G-2426-SM-7DUGM
60-69 years, male
1056.1
GTEX-WYJK-1826-SM-4ONDM
50-59 years, male
1055.7
GTEX-XBEC-0126-SM-4GIDT
50-59 years, male
1053.0
GTEX-144GM-0426-SM-5Q5C8
20-29 years, male
1052.9
GTEX-S4Z8-2126-SM-4AD5H
60-69 years, male
1051.5
GTEX-13112-0226-SM-5P9IV
50-59 years, male
1050.0
GTEX-14C5O-2326-SM-73KYU
60-69 years, male
1049.3
GTEX-QLQ7-1426-SM-4R1JX
60-69 years, male
1048.3
GTEX-YJ89-0626-SM-4TT3Z
60-69 years, male
1048.3
GTEX-14PJ4-1726-SM-664OC
60-69 years, male
1045.1
GTEX-17HGU-0726-SM-7DUFN
50-59 years, male
1042.7
GTEX-T5JC-0726-SM-4DM55
20-29 years, male
1042.1
GTEX-11NSD-1026-SM-5N9BE
20-29 years, male
1039.1
GTEX-R55D-0126-SM-48FEL
50-59 years, male
1037.8
GTEX-YEC4-1526-SM-4W1YU
40-49 years, male
1036.6
GTEX-XGQ4-2026-SM-4AT6G
50-59 years, male
1032.1
GTEX-18D9B-2726-SM-72D73
60-69 years, male
1030.1
GTEX-18A6Q-2626-SM-7KFT3
60-69 years, male
1029.7
GTEX-OIZH-2126-SM-3NB1P
50-59 years, male
1027.8
GTEX-Y5V6-1726-SM-4VDSZ
60-69 years, male
1027.1
GTEX-S4Q7-1226-SM-4AD5I
20-29 years, male
1025.9
GTEX-1B8KE-1426-SM-7EWEL
50-59 years, male
1025.5
GTEX-Q2AH-1526-SM-48TZG
40-49 years, male
1025.5
GTEX-QEG4-0126-SM-48TZE
30-39 years, male
1024.4
GTEX-139TT-2226-SM-5LZWO
60-69 years, male
1024.0
GTEX-12BJ1-1326-SM-5BC5P
60-69 years, male
1023.0
GTEX-1AX9I-1826-SM-72D5I
60-69 years, male
1022.9
GTEX-S95S-1126-SM-4B64E
60-69 years, male
1020.6
GTEX-OXRL-2126-SM-3NM98
50-59 years, male
1020.3
GTEX-144GL-0726-SM-5LU4P
50-59 years, male
1019.3
GTEX-P4QS-2126-SM-3NMCF
60-69 years, male
1018.4
GTEX-ZTX8-1126-SM-51MRM
20-29 years, male
1018.1
GTEX-1399R-1626-SM-5P9GG
30-39 years, male
1014.5
GTEX-14C39-0526-SM-664OF
40-49 years, male
1008.7
GTEX-ZA64-1626-SM-5CVME
20-29 years, male
1008.0
GTEX-QV44-1726-SM-4R1KG
50-59 years, male
1007.8
GTEX-YB5E-1926-SM-5IFIG
40-49 years, male
1005.9
GTEX-R55E-0726-SM-48FCZ
20-29 years, male
1004.2
GTEX-R55C-1426-SM-48FED
40-49 years, male
1003.6
GTEX-14A6H-2326-SM-5Q5B5
30-39 years, male
1002.1
GTEX-145MO-0126-SM-5S2QU
60-69 years, male
1002.0
GTEX-15RIE-0626-SM-6M47G
60-69 years, male
1001.4
GTEX-15SHV-1926-SM-6LPI1
60-69 years, male
997.9
GTEX-11TT1-2226-SM-5GU6B
20-29 years, male
996.8
GTEX-13N1W-2626-SM-5IJEP
70-79 years, male
995.0
GTEX-1A3MV-0726-SM-72D5M
30-39 years, male
994.4
GTEX-13NYB-2226-SM-5MR58
40-49 years, male
993.9
GTEX-12ZZY-0126-SM-5LZV2
60-69 years, male
991.1
GTEX-144GN-1626-SM-5Q5BU
50-59 years, male
990.1
GTEX-18A7B-1526-SM-7KFTH
70-79 years, male
989.7
GTEX-14E1K-1926-SM-73KWS
50-59 years, male
988.1
GTEX-13OW6-0126-SM-5IJGM
50-59 years, male
987.4
GTEX-U8T8-1126-SM-4DXUE
60-69 years, male
982.0
GTEX-Q2AI-1226-SM-48U14
40-49 years, male
978.1
GTEX-U3ZM-1626-SM-4DXSK
40-49 years, male
976.7
GTEX-RN64-2326-SM-48FDW
50-59 years, male
975.7
GTEX-WZTO-0326-SM-4PQYZ
40-49 years, male
974.3
GTEX-11EQ8-1426-SM-5EGJR
60-69 years, male
971.6
GTEX-XPT6-1626-SM-4B655
20-29 years, male
969.2
GTEX-147F4-0626-SM-5LUAK
50-59 years, male
968.7
GTEX-15G19-1326-SM-6LPIR
60-69 years, male
967.8
GTEX-13OVK-2226-SM-6LPJY
40-49 years, male
965.4
GTEX-15RIF-1826-SM-6M469
20-29 years, male
964.3
GTEX-T6MN-2026-SM-4DM7L
50-59 years, male
961.5
GTEX-XMK1-2026-SM-4B65K
40-49 years, male
961.3
GTEX-QMRM-1526-SM-4R1K6
50-59 years, male
961.0
GTEX-17MFQ-0826-SM-793C8
20-29 years, male
959.7
GTEX-111CU-1726-SM-5EGHM
50-59 years, male
959.0
GTEX-17HHY-0726-SM-7IGM6
50-59 years, male
959.0
GTEX-Y111-2426-SM-4TT23
50-59 years, male
959.0
GTEX-17HII-0126-SM-7KFSS
50-59 years, male
958.5
GTEX-QV31-1126-SM-4R1K4
30-39 years, male
958.4
GTEX-XPVG-2226-SM-4B65U
50-59 years, male
957.7
GTEX-ZDYS-1326-SM-5IJFF
50-59 years, male
957.3
GTEX-OOBJ-2126-SM-3NB1N
60-69 years, male
956.4
GTEX-17HHE-1226-SM-793C6
50-59 years, male
955.2
GTEX-145MH-2326-SM-5O9AW
50-59 years, male
951.9
GTEX-1CB4J-1626-SM-79OOK
60-69 years, male
950.7
GTEX-14DAR-1126-SM-793AT
50-59 years, male
950.2
GTEX-11NV4-1726-SM-5N9FC
60-69 years, male
948.3
GTEX-13N2G-0126-SM-5N9DV
60-69 years, male
948.0
GTEX-U4B1-1526-SM-4DXSL
40-49 years, male
947.2
GTEX-11O72-0726-SM-5P9GO
40-49 years, male
946.8
GTEX-1CB4I-0926-SM-7MKFY
50-59 years, male
946.1
GTEX-S7PM-0626-SM-4AD4Q
60-69 years, male
945.8
GTEX-WHSE-0426-SM-4M1XO
20-29 years, male
944.8
GTEX-13O1R-0726-SM-5IJEI
60-69 years, male
943.3
GTEX-1CAMR-1526-SM-79OLP
20-29 years, male
943.3
GTEX-145MF-1726-SM-5LU9H
50-59 years, male
942.9
GTEX-Y8E4-2226-SM-5LU94
40-49 years, male
941.1
GTEX-147JS-0126-SM-5S2TW
60-69 years, male
939.0
GTEX-13OVH-0726-SM-5N9BU
70-79 years, male
937.9
GTEX-WOFM-1126-SM-4OOSB
20-29 years, male
937.7
GTEX-12WSH-0326-SM-5GCNH
50-59 years, male
937.5
GTEX-REY6-0126-SM-48FDT
60-69 years, male
936.9
GTEX-11DXY-0226-SM-5H123
60-69 years, male
936.4
GTEX-11P7K-1026-SM-5HL6Y
30-39 years, male
934.6
GTEX-15SKB-0326-SM-6M477
60-69 years, male
932.2
GTEX-QEG5-0126-SM-4R1JR
20-29 years, male
930.5
GTEX-PW2O-1426-SM-48TCD
20-29 years, male
929.3
GTEX-NFK9-0126-SM-3LK5H
40-49 years, male
929.0
GTEX-RWSA-2426-SM-47JXR
40-49 years, male
927.1
GTEX-SNOS-1126-SM-4DM67
40-49 years, male
925.0
GTEX-183FY-1926-SM-7KFRI
20-29 years, male
922.7
GTEX-X5EB-2026-SM-4E3KA
40-49 years, male
922.2
GTEX-11GS4-2026-SM-5N9CP
60-69 years, male
920.4
GTEX-1B996-2326-SM-731EC
60-69 years, male
919.2
GTEX-VJYA-1426-SM-4KL1Y
60-69 years, male
919.0
GTEX-111VG-1926-SM-5GIDO
60-69 years, male
918.3
GTEX-OIZI-0126-SM-3NB13
40-49 years, male
918.3
GTEX-ZQUD-2026-SM-51MSM
30-39 years, male
918.3
GTEX-145LT-0426-SM-5LUAP
40-49 years, male
916.8
GTEX-11DXZ-2126-SM-59881
50-59 years, male
916.7
GTEX-14PJ2-2026-SM-6AJAQ
50-59 years, male
915.9
GTEX-V955-1926-SM-4KL1L
60-69 years, male
915.8
GTEX-ZT9X-1426-SM-5DUX1
40-49 years, male
915.7
GTEX-139T6-1226-SM-5IFFC
50-59 years, male
914.5
GTEX-117XS-2026-SM-5GID1
60-69 years, male
914.3
GTEX-ZUA1-2726-SM-59HLJ
40-49 years, male
913.8
GTEX-ZV7C-2026-SM-5NQ8F
50-59 years, male
913.2
GTEX-16MTA-1126-SM-6LPJA
50-59 years, male
912.0
GTEX-17GQL-1726-SM-718B8
60-69 years, male
911.5
GTEX-1C4CL-1926-SM-731DY
30-39 years, male
908.9
GTEX-1F6IF-0526-SM-7MKHD
60-69 years, male
907.8
GTEX-ZT9W-2226-SM-57WFU
50-59 years, male
907.5
GTEX-U8XE-0126-SM-4E3I3
30-39 years, male
906.1
GTEX-13FHP-2826-SM-5IJFW
50-59 years, male
903.7
GTEX-1EKGG-2726-SM-79OOD
60-69 years, male
902.7
GTEX-Y9LG-1726-SM-4VBQE
30-39 years, male
901.5
GTEX-XLM4-1526-SM-4AT6D
60-69 years, male
901.2
GTEX-ZYFC-0126-SM-5GIEH
50-59 years, male
898.3
GTEX-TKQ2-1526-SM-4DXUN
50-59 years, male
894.5
GTEX-RUSQ-2126-SM-47JXK
50-59 years, male
893.6
GTEX-14ABY-0626-SM-5Q5C9
50-59 years, male
892.9
GTEX-WH7G-1926-SM-4LVMM
40-49 years, male
892.3
GTEX-1212Z-0326-SM-5FQSJ
60-69 years, male
891.3
GTEX-YEC3-1726-SM-5IFIK
50-59 years, male
890.3
GTEX-12WSL-2326-SM-5DUXQ
50-59 years, male
889.4
GTEX-132QS-1226-SM-5P9GD
60-69 years, male
888.4
GTEX-ZVTK-0126-SM-57WDG
20-29 years, male
887.1
GTEX-14PJN-1626-SM-68727
30-39 years, male
886.2
GTEX-ZVZP-2226-SM-57WBF
50-59 years, male
886.0
GTEX-18A67-1826-SM-7KFT7
50-59 years, male
884.8
GTEX-14E7W-0726-SM-664OK
70-79 years, male
881.6
GTEX-14A5H-0626-SM-5TDCO
60-69 years, male
876.5
GTEX-WHSB-2126-SM-4M1XF
50-59 years, male
875.5
GTEX-12696-0226-SM-5EGL3
60-69 years, male
875.1
GTEX-V1D1-2126-SM-4JBH4
50-59 years, male
871.6
GTEX-13OVL-0426-SM-5IFG6
50-59 years, male
869.2
GTEX-ZYT6-2726-SM-5GICP
30-39 years, male
864.9
GTEX-11ONC-2226-SM-5HL6D
60-69 years, male
864.3
GTEX-1BAJH-0726-SM-7IGMF
70-79 years, male
862.9
GTEX-ZLFU-2026-SM-4WWG2
40-49 years, male
862.8
GTEX-14B4R-1026-SM-5TDDS
30-39 years, male
862.1
GTEX-1E2YA-2826-SM-7EPIN
50-59 years, male
860.8
GTEX-1B8SF-2926-SM-731DL
60-69 years, male
860.0
GTEX-131XE-0426-SM-5IJF4
50-59 years, male
857.2
GTEX-11NUK-2626-SM-5A5MB
50-59 years, male
855.6
GTEX-16XZZ-0626-SM-6M47T
60-69 years, male
851.1
GTEX-11EQ9-1926-SM-5PNVV
30-39 years, male
847.2
GTEX-X3Y1-2626-SM-4PQZI
40-49 years, male
843.3
GTEX-ZAB4-0126-SM-5CVMG
40-49 years, male
841.9
GTEX-14PHX-1426-SM-69LPP
50-59 years, male
839.2
GTEX-ZZ64-1126-SM-5GZXY
20-29 years, male
838.8
GTEX-111YS-2026-SM-5EGGL
60-69 years, male
838.6
GTEX-14PJ3-1626-SM-664O5
50-59 years, male
836.6
GTEX-1399T-1526-SM-5P9J6
60-69 years, male
834.4
GTEX-11WQC-2326-SM-5EQKE
60-69 years, male
834.2
GTEX-11OF3-1826-SM-5987N
60-69 years, male
832.7
GTEX-15RJ7-1826-SM-7KUMJ
40-49 years, male
820.2
GTEX-13OW5-2526-SM-5L3I1
60-69 years, male
819.2
GTEX-XAJ8-1326-SM-47JYT
40-49 years, male
818.4
GTEX-12WSI-2126-SM-5GCMV
50-59 years, male
818.3
GTEX-1339X-1926-SM-5PNVP
40-49 years, male
816.5
GTEX-N7MS-0126-SM-3TW8O
60-69 years, male
814.9
GTEX-1A3MX-1026-SM-731F8
60-69 years, male
812.7
GTEX-1GMRU-0626-SM-7MKH2
50-59 years, male
812.0
GTEX-Z93S-1726-SM-5HL8G
50-59 years, male
811.9
GTEX-11LCK-2326-SM-5HL53
30-39 years, male
809.4
GTEX-14753-0626-SM-5Q5CY
60-69 years, male
809.3
GTEX-1E1VI-1026-SM-7MKGQ
50-59 years, male
806.3
GTEX-ZAB5-2426-SM-5CVMW
50-59 years, male
804.9
GTEX-12C56-1426-SM-5FQSW
20-29 years, male
800.4
GTEX-13OW8-0526-SM-5KM24
60-69 years, male
798.6
GTEX-11ZUS-2726-SM-5FQUA
30-39 years, male
793.4
GTEX-147GR-0626-SM-5S2PK
60-69 years, male
790.5
GTEX-1AX8Z-2826-SM-73KTZ
60-69 years, male
788.9
GTEX-PLZ6-1226-SM-3P5ZS
30-39 years, male
787.3
GTEX-WVJS-2726-SM-4MVNU
50-59 years, male
780.7
GTEX-13FLW-2126-SM-5N9FD
50-59 years, male
776.6
GTEX-1EU9M-1626-SM-7EWF2
60-69 years, male
775.8
GTEX-117YW-1526-SM-5EGGP
50-59 years, male
774.0
GTEX-16Z82-2726-SM-7KULV
60-69 years, male
771.7
GTEX-18QFQ-1926-SM-72D5H
30-39 years, male
766.0
GTEX-Y3I4-2026-SM-4TT6Z
50-59 years, male
762.2
GTEX-17F9E-1626-SM-7EWDB
50-59 years, male
757.0
GTEX-1C64O-2126-SM-7IGPB
60-69 years, male
755.4
GTEX-13111-1526-SM-5EGJX
50-59 years, male
751.3
GTEX-17MF6-2726-SM-7IGMZ
40-49 years, male
746.0
GTEX-16AAH-1626-SM-7EWE2
50-59 years, male
744.8
GTEX-ZPU1-2126-SM-57WED
40-49 years, male
735.9
GTEX-1C6VQ-1426-SM-73KTW
50-59 years, male
726.5
GTEX-111FC-1926-SM-5GZYC
60-69 years, male
724.6
GTEX-1C6VR-1526-SM-7MKG9
60-69 years, male
723.9
GTEX-11P82-1526-SM-5BC5M
20-29 years, male
723.8
GTEX-148VJ-2526-SM-5TDEI
70-79 years, male
711.6
GTEX-1C64N-2826-SM-7IGP5
50-59 years, male
697.6
GTEX-12WSM-1326-SM-5GCP9
60-69 years, male
677.7
GTEX-13QJ3-0226-SM-5S2PU
50-59 years, male
670.9
GTEX-1C6VS-1126-SM-7EWEW
60-69 years, male
659.4
GTEX-17F97-2726-SM-7IGQ4
50-59 years, male
647.4
GTEX-11TUW-2226-SM-5EQL9
60-69 years, male
632.6
GTEX-NPJ8-1226-SM-3MJHM
40-49 years, male
572.7
GTEX-1B8KZ-1826-SM-73KVJ
50-59 years, male
555.8
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TESTIS - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.