We use cookies to enhance the usability of our website. If you continue, we'll assume that you are happy to receive all cookies. More information. Don't show this again.
On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression (NX) levels for human tissues was created by combining the data from these three transcriptomics datasets (HPA, GTEx and FANTOM5).
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
Antibody staining in the annotated cell types in the current human tissue is reported as not detected, low, medium, or high. This score is based on the staining intensity and fraction of stained cells.
The RNA-seq details section shows detailed information about the individual samples used for the transcript profiling and results of the RNA-seq analysis.
Information about each individual sample is listed below, including gender, age, a tissue section image and estimated fractions of cell types. pTPM (transcripts per million) values give a quantification of the gene abundance which is comparable between different genes and samples.
Cells in seminiferous ducts: 65 Other cell types: 35
Male, age 34
Testis sample 278 pTPM: 651.8
Cell types%
Cells in seminiferous ducts: 75 Other cell types: 25
Male, age 62
Testis sample 279 pTPM: 574.8
Cell types%
Cells in seminiferous ducts: 70 Other cell types: 30
Male, age 34
Testis sample 280 pTPM: 1163.4
Cell types%
Cells in seminiferous ducts: 60 Other cell types: 40
Male, age 26
Testis sample 281 pTPM: 517.7
Cell types%
Cells in seminiferous ducts: 55 Other cell types: 45
Male, age 50
Testis sample 282 pTPM: 401.5
Cell types%
Cells in seminiferous ducts: 85 Other cell types: 15
Male, age 37
Testis sample 283 pTPM: 689.4
Cell types%
Cells in seminiferous ducts: 35 Other cell types: 65
Male, age 51
Testis sample 349 pTPM: 1022.2
Cell types%
Cells in seminiferous ducts: 65 Other cell types: 35
Male, age 34
Testis sample 354 pTPM: 1081.2
Cell types%
Cells in seminiferous ducts: 70 Other cell types: 30
Male, age 26
Testis sample 371 pTPM: 1172
Cell types%
Cells in seminiferous ducts: 75 Other cell types: 25
TESTIS - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as average pTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. pTPM values of the individual samples are presented next to the box plot.
Max pTPM: 3576.9
Min pTPM: 159.9
Std pTPM: 355.7
Median pTPM: 603.2
GTEX-NPJ8-1226-SM-3MJHM
40-49 years, male
3576.9
GTEX-1C6VQ-1426-SM-73KTW
50-59 years, male
2173.5
GTEX-148VJ-2526-SM-5TDEI
70-79 years, male
2131.9
GTEX-17F97-2726-SM-7IGQ4
50-59 years, male
2028.9
GTEX-11DXY-0226-SM-5H123
60-69 years, male
1604.7
GTEX-PLZ6-1226-SM-3P5ZS
30-39 years, male
1581.9
GTEX-1AX8Z-2826-SM-73KTZ
60-69 years, male
1565.4
GTEX-Y3I4-2026-SM-4TT6Z
50-59 years, male
1559.2
GTEX-1C6VS-1126-SM-7EWEW
60-69 years, male
1554.6
GTEX-18A67-1826-SM-7KFT7
50-59 years, male
1488.9
GTEX-1399T-1526-SM-5P9J6
60-69 years, male
1467.8
GTEX-12WSM-1326-SM-5GCP9
60-69 years, male
1461.5
GTEX-13QJ3-0226-SM-5S2PU
50-59 years, male
1337.5
GTEX-147GR-0626-SM-5S2PK
60-69 years, male
1330.1
GTEX-1CB4J-1626-SM-79OOK
60-69 years, male
1301.1
GTEX-S4Z8-2126-SM-4AD5H
60-69 years, male
1288.9
GTEX-1B8KZ-1826-SM-73KVJ
50-59 years, male
1281.7
GTEX-ZLFU-2026-SM-4WWG2
40-49 years, male
1277.0
GTEX-145LT-0426-SM-5LUAP
40-49 years, male
1268.2
GTEX-13OW8-0526-SM-5KM24
60-69 years, male
1237.2
GTEX-147F4-0626-SM-5LUAK
50-59 years, male
1166.4
GTEX-14A6H-2326-SM-5Q5B5
30-39 years, male
1148.2
GTEX-YJ89-0626-SM-4TT3Z
60-69 years, male
1118.0
GTEX-1E2YA-2826-SM-7EPIN
50-59 years, male
1099.0
GTEX-1399R-1626-SM-5P9GG
30-39 years, male
1089.5
GTEX-WY7C-2226-SM-4ONCS
50-59 years, male
1085.3
GTEX-WVJS-2726-SM-4MVNU
50-59 years, male
1067.9
GTEX-QEG5-0126-SM-4R1JR
20-29 years, male
1065.7
GTEX-145MF-1726-SM-5LU9H
50-59 years, male
1063.1
GTEX-NFK9-0126-SM-3LK5H
40-49 years, male
1029.6
GTEX-ZAB4-0126-SM-5CVMG
40-49 years, male
1016.9
GTEX-OIZI-0126-SM-3NB13
40-49 years, male
989.8
GTEX-13OW6-0126-SM-5IJGM
50-59 years, male
984.7
GTEX-WHSB-2126-SM-4M1XF
50-59 years, male
966.3
GTEX-12BJ1-1326-SM-5BC5P
60-69 years, male
963.2
GTEX-15RJ7-1826-SM-7KUMJ
40-49 years, male
951.3
GTEX-PW2O-1426-SM-48TCD
20-29 years, male
948.6
GTEX-11TT1-2226-SM-5GU6B
20-29 years, male
948.3
GTEX-117YX-2026-SM-5GIEF
50-59 years, male
876.1
GTEX-Y8E4-2226-SM-5LU94
40-49 years, male
870.0
GTEX-1BAJH-0726-SM-7IGMF
70-79 years, male
864.7
GTEX-15SKB-0326-SM-6M477
60-69 years, male
859.7
GTEX-11ONC-2226-SM-5HL6D
60-69 years, male
856.4
GTEX-14B4R-1026-SM-5TDDS
30-39 years, male
847.3
GTEX-S3XE-1526-SM-4AD5A
50-59 years, male
845.3
GTEX-QV44-1726-SM-4R1KG
50-59 years, male
836.6
GTEX-14A5H-0626-SM-5TDCO
60-69 years, male
836.2
GTEX-YF7O-2026-SM-4W1YE
50-59 years, male
824.5
GTEX-O5YT-2126-SM-3MJGD
20-29 years, male
823.4
GTEX-WFG8-1926-SM-4LVM1
20-29 years, male
822.9
GTEX-U3ZH-1526-SM-4DXV1
30-39 years, male
822.8
GTEX-OIZH-2126-SM-3NB1P
50-59 years, male
821.7
GTEX-ZDTT-2026-SM-5K7TY
60-69 years, male
821.3
GTEX-T5JC-0726-SM-4DM55
20-29 years, male
815.3
GTEX-OOBJ-2126-SM-3NB1N
60-69 years, male
812.2
GTEX-V955-1926-SM-4KL1L
60-69 years, male
801.0
GTEX-Y9LG-1726-SM-4VBQE
30-39 years, male
793.1
GTEX-VJYA-1426-SM-4KL1Y
60-69 years, male
791.3
GTEX-P4QS-2126-SM-3NMCF
60-69 years, male
785.5
GTEX-1339X-1926-SM-5PNVP
40-49 years, male
781.1
GTEX-17HGU-0726-SM-7DUFN
50-59 years, male
779.0
GTEX-1AYCT-1026-SM-79ONR
50-59 years, male
777.6
GTEX-S33H-0126-SM-4AD62
30-39 years, male
777.3
GTEX-U8T8-1126-SM-4DXUE
60-69 years, male
774.2
GTEX-R55C-1426-SM-48FED
40-49 years, male
763.6
GTEX-ZQUD-2026-SM-51MSM
30-39 years, male
761.9
GTEX-RN64-2326-SM-48FDW
50-59 years, male
761.1
GTEX-QLQ7-1426-SM-4R1JX
60-69 years, male
758.3
GTEX-144GN-1626-SM-5Q5BU
50-59 years, male
754.7
GTEX-14PHX-1426-SM-69LPP
50-59 years, male
740.4
GTEX-YEC4-1526-SM-4W1YU
40-49 years, male
736.2
GTEX-14PJ4-1726-SM-664OC
60-69 years, male
734.9
GTEX-13N2G-0126-SM-5N9DV
60-69 years, male
725.9
GTEX-1CB4I-0926-SM-7MKFY
50-59 years, male
723.7
GTEX-14C38-0126-SM-5YY9V
50-59 years, male
722.7
GTEX-PLZ5-1526-SM-3P5ZX
50-59 years, male
721.2
GTEX-QMRM-1526-SM-4R1K6
50-59 years, male
718.3
GTEX-11P7K-1026-SM-5HL6Y
30-39 years, male
713.5
GTEX-WFON-2026-SM-4LVMW
40-49 years, male
712.4
GTEX-RWSA-2426-SM-47JXR
40-49 years, male
705.1
GTEX-ZTX8-1126-SM-51MRM
20-29 years, male
704.9
GTEX-13NYB-2226-SM-5MR58
40-49 years, male
701.0
GTEX-X261-2326-SM-4PQYU
50-59 years, male
696.5
GTEX-YFCO-1726-SM-4W21S
40-49 years, male
695.1
GTEX-XGQ4-2026-SM-4AT6G
50-59 years, male
694.4
GTEX-1AX9K-1826-SM-731CY
50-59 years, male
693.1
GTEX-16MTA-1126-SM-6LPJA
50-59 years, male
692.9
GTEX-12WSI-2126-SM-5GCMV
50-59 years, male
684.3
GTEX-17MF6-2726-SM-7IGMZ
40-49 years, male
682.2
GTEX-183FY-1926-SM-7KFRI
20-29 years, male
681.6
GTEX-XBED-2026-SM-4AT5D
60-69 years, male
681.4
GTEX-17KNJ-1126-SM-7KFT6
50-59 years, male
678.3
GTEX-XPT6-1626-SM-4B655
20-29 years, male
676.3
GTEX-YB5E-1926-SM-5IFIG
40-49 years, male
673.8
GTEX-XMK1-2026-SM-4B65K
40-49 years, male
673.0
GTEX-14PJN-1626-SM-68727
30-39 years, male
672.0
GTEX-139TT-2226-SM-5LZWO
60-69 years, male
671.5
GTEX-TKQ1-0926-SM-4DXU2
30-39 years, male
670.6
GTEX-1B996-2326-SM-731EC
60-69 years, male
669.6
GTEX-14E7W-0726-SM-664OK
70-79 years, male
668.7
GTEX-U3ZM-1626-SM-4DXSK
40-49 years, male
666.7
GTEX-XPVG-2226-SM-4B65U
50-59 years, male
662.3
GTEX-14753-0626-SM-5Q5CY
60-69 years, male
654.8
GTEX-Y5V6-1726-SM-4VDSZ
60-69 years, male
653.4
GTEX-X3Y1-2626-SM-4PQZI
40-49 years, male
650.6
GTEX-13OVK-2226-SM-6LPJY
40-49 years, male
649.6
GTEX-13O61-2026-SM-5J2M6
60-69 years, male
647.5
GTEX-S7SE-0326-SM-4AT5Q
50-59 years, male
647.3
GTEX-XLM4-1526-SM-4AT6D
60-69 years, male
647.2
GTEX-14E6E-1026-SM-664N9
30-39 years, male
645.3
GTEX-15RIE-0626-SM-6M47G
60-69 years, male
642.9
GTEX-RM2N-1326-SM-48FCW
50-59 years, male
641.6
GTEX-131XE-0426-SM-5IJF4
50-59 years, male
641.2
GTEX-WVLH-2626-SM-4MVNV
50-59 years, male
639.5
GTEX-11O72-0726-SM-5P9GO
40-49 years, male
636.5
GTEX-U4B1-1526-SM-4DXSL
40-49 years, male
636.0
GTEX-13FTW-1326-SM-5LZZD
40-49 years, male
632.9
GTEX-13OVH-0726-SM-5N9BU
70-79 years, male
630.2
GTEX-R55D-0126-SM-48FEL
50-59 years, male
625.5
GTEX-18A7B-1526-SM-7KFTH
70-79 years, male
623.6
GTEX-ZT9X-1426-SM-5DUX1
40-49 years, male
622.7
GTEX-1CAMR-1526-SM-79OLP
20-29 years, male
621.5
GTEX-ZV7C-2026-SM-5NQ8F
50-59 years, male
617.9
GTEX-15SHV-1926-SM-6LPI1
60-69 years, male
614.4
GTEX-WHSE-0426-SM-4M1XO
20-29 years, male
612.8
GTEX-1A3MX-1026-SM-731F8
60-69 years, male
610.6
GTEX-139T6-1226-SM-5IFFC
50-59 years, male
610.4
GTEX-16Z82-2726-SM-7KULV
60-69 years, male
610.0
GTEX-WH7G-1926-SM-4LVMM
40-49 years, male
605.6
GTEX-Q2AH-1526-SM-48TZG
40-49 years, male
603.2
GTEX-14DAR-1126-SM-793AT
50-59 years, male
602.6
GTEX-17F9E-1626-SM-7EWDB
50-59 years, male
601.9
GTEX-147JS-0126-SM-5S2TW
60-69 years, male
600.2
GTEX-15RIF-1826-SM-6M469
20-29 years, male
598.7
GTEX-18D9B-2726-SM-72D73
60-69 years, male
597.7
GTEX-11EI6-2226-SM-5EGJM
60-69 years, male
597.4
GTEX-16XZZ-0626-SM-6M47T
60-69 years, male
595.8
GTEX-1F6IF-0526-SM-7MKHD
60-69 years, male
594.6
GTEX-16YQH-1826-SM-6LPJW
30-39 years, male
593.0
GTEX-RUSQ-2126-SM-47JXK
50-59 years, male
591.1
GTEX-T8EM-0126-SM-4DM5R
40-49 years, male
589.2
GTEX-13NZB-2026-SM-5MR4M
30-39 years, male
586.7
GTEX-QLQW-1026-SM-447A9
30-39 years, male
581.6
GTEX-11P82-1526-SM-5BC5M
20-29 years, male
579.5
GTEX-ZZ64-1126-SM-5GZXY
20-29 years, male
577.4
GTEX-ZYFC-0126-SM-5GIEH
50-59 years, male
568.7
GTEX-ZVZP-2226-SM-57WBF
50-59 years, male
568.2
GTEX-1212Z-0326-SM-5FQSJ
60-69 years, male
566.4
GTEX-14BIL-2226-SM-73KWF
50-59 years, male
565.8
GTEX-14E1K-1926-SM-73KWS
50-59 years, male
561.5
GTEX-ZVTK-0126-SM-57WDG
20-29 years, male
561.2
GTEX-11NSD-1026-SM-5N9BE
20-29 years, male
560.4
GTEX-11GS4-2026-SM-5N9CP
60-69 years, male
555.0
GTEX-ZUA1-2726-SM-59HLJ
40-49 years, male
552.4
GTEX-132QS-1226-SM-5P9GD
60-69 years, male
552.3
GTEX-SNMC-1026-SM-4DM7K
20-29 years, male
551.9
GTEX-12C56-1426-SM-5FQSW
20-29 years, male
549.4
GTEX-144GL-0726-SM-5LU4P
50-59 years, male
547.9
GTEX-S95S-1126-SM-4B64E
60-69 years, male
544.5
GTEX-WYJK-1826-SM-4ONDM
50-59 years, male
542.7
GTEX-TKQ2-1526-SM-4DXUN
50-59 years, male
541.8
GTEX-R55E-0726-SM-48FCZ
20-29 years, male
541.6
GTEX-ZA64-1626-SM-5CVME
20-29 years, male
538.8
GTEX-1B8KE-1426-SM-7EWEL
50-59 years, male
538.6
GTEX-ZAB5-2426-SM-5CVMW
50-59 years, male
538.2
GTEX-U8XE-0126-SM-4E3I3
30-39 years, male
536.1
GTEX-1AX9I-1826-SM-72D5I
60-69 years, male
535.9
GTEX-ZDYS-1326-SM-5IJFF
50-59 years, male
533.5
GTEX-13111-1526-SM-5EGJX
50-59 years, male
530.8
GTEX-OXRL-2126-SM-3NM98
50-59 years, male
529.3
GTEX-18QFQ-1926-SM-72D5H
30-39 years, male
528.1
GTEX-1E1VI-1026-SM-7MKGQ
50-59 years, male
528.0
GTEX-18A66-0726-SM-72D77
60-69 years, male
527.7
GTEX-18465-1526-SM-7KFTV
50-59 years, male
524.6
GTEX-S4Q7-1226-SM-4AD5I
20-29 years, male
523.4
GTEX-111CU-1726-SM-5EGHM
50-59 years, male
522.3
GTEX-139TS-1726-SM-5IJG5
60-69 years, male
520.1
GTEX-ZT9W-2226-SM-57WFU
50-59 years, male
516.2
GTEX-11DXZ-2126-SM-59881
50-59 years, male
516.1
GTEX-14C39-0526-SM-664OF
40-49 years, male
515.5
GTEX-13FLW-2126-SM-5N9FD
50-59 years, male
515.4
GTEX-XBEC-0126-SM-4GIDT
50-59 years, male
514.4
GTEX-SNOS-1126-SM-4DM67
40-49 years, male
512.2
GTEX-V1D1-2126-SM-4JBH4
50-59 years, male
509.6
GTEX-13O1R-0726-SM-5IJEI
60-69 years, male
508.7
GTEX-REY6-0126-SM-48FDT
60-69 years, male
506.7
GTEX-14PJ3-1626-SM-664O5
50-59 years, male
506.2
GTEX-145MH-2326-SM-5O9AW
50-59 years, male
504.7
GTEX-1C64O-2126-SM-7IGPB
60-69 years, male
504.4
GTEX-117XS-2026-SM-5GID1
60-69 years, male
502.2
GTEX-15G19-1326-SM-6LPIR
60-69 years, male
501.1
GTEX-11NUK-2626-SM-5A5MB
50-59 years, male
499.0
GTEX-1AX9J-1226-SM-72D6P
60-69 years, male
498.9
GTEX-QV31-1126-SM-4R1K4
30-39 years, male
496.8
GTEX-145MO-0126-SM-5S2QU
60-69 years, male
495.7
GTEX-12WSH-0326-SM-5GCNH
50-59 years, male
491.4
GTEX-ZPU1-2126-SM-57WED
40-49 years, male
491.0
GTEX-Z93S-1726-SM-5HL8G
50-59 years, male
490.2
GTEX-1A3MV-0726-SM-72D5M
30-39 years, male
488.8
GTEX-13O21-1226-SM-5J2MK
50-59 years, male
488.5
GTEX-12ZZY-0126-SM-5LZV2
60-69 years, male
486.3
GTEX-17HHE-1226-SM-793C6
50-59 years, male
485.1
GTEX-13N1W-2626-SM-5IJEP
70-79 years, male
482.4
GTEX-18A6Q-2626-SM-7KFT3
60-69 years, male
477.2
GTEX-1B8SF-2926-SM-731DL
60-69 years, male
477.2
GTEX-OHPM-2126-SM-3LK75
50-59 years, male
474.4
GTEX-14C5O-2326-SM-73KYU
60-69 years, male
473.6
GTEX-11OF3-1826-SM-5987N
60-69 years, male
473.5
GTEX-12696-0226-SM-5EGL3
60-69 years, male
472.7
GTEX-12WSL-2326-SM-5DUXQ
50-59 years, male
470.5
GTEX-17GQL-1726-SM-718B8
60-69 years, male
465.4
GTEX-T6MN-2026-SM-4DM7L
50-59 years, male
464.8
GTEX-ZYT6-2726-SM-5GICP
30-39 years, male
464.5
GTEX-13OW5-2526-SM-5L3I1
60-69 years, male
460.2
GTEX-17HHY-0726-SM-7IGM6
50-59 years, male
458.7
GTEX-1GMRU-0626-SM-7MKH2
50-59 years, male
458.1
GTEX-11EQ8-1426-SM-5EGJR
60-69 years, male
456.1
GTEX-17HII-0126-SM-7KFSS
50-59 years, male
454.9
GTEX-ZTSS-1526-SM-51MTC
40-49 years, male
453.5
GTEX-WOFM-1126-SM-4OOSB
20-29 years, male
448.5
GTEX-11EQ9-1926-SM-5PNVV
30-39 years, male
446.2
GTEX-UPJH-0126-SM-4IHLL
50-59 years, male
445.4
GTEX-111YS-2026-SM-5EGGL
60-69 years, male
441.8
GTEX-Q2AI-1226-SM-48U14
40-49 years, male
441.3
GTEX-144GM-0426-SM-5Q5C8
20-29 years, male
440.0
GTEX-XAJ8-1326-SM-47JYT
40-49 years, male
435.3
GTEX-OOBK-2126-SM-3LK5T
40-49 years, male
432.2
GTEX-111FC-1926-SM-5GZYC
60-69 years, male
431.7
GTEX-13VXU-0726-SM-5J2O7
50-59 years, male
423.7
GTEX-11LCK-2326-SM-5HL53
30-39 years, male
421.7
GTEX-14ABY-0626-SM-5Q5C9
50-59 years, male
418.3
GTEX-1EU9M-1626-SM-7EWF2
60-69 years, male
416.0
GTEX-13NZA-2526-SM-5IJFX
60-69 years, male
414.4
GTEX-16AAH-1626-SM-7EWE2
50-59 years, male
414.3
GTEX-14PJ2-2026-SM-6AJAQ
50-59 years, male
407.8
GTEX-YEC3-1726-SM-5IFIK
50-59 years, male
401.8
GTEX-N7MS-0126-SM-3TW8O
60-69 years, male
395.3
GTEX-Y111-2426-SM-4TT23
50-59 years, male
394.1
GTEX-13OVL-0426-SM-5IFG6
50-59 years, male
390.9
GTEX-1CB4G-2426-SM-7DUGM
60-69 years, male
379.7
GTEX-1C4CL-1926-SM-731DY
30-39 years, male
379.7
GTEX-X5EB-2026-SM-4E3KA
40-49 years, male
375.4
GTEX-WZTO-0326-SM-4PQYZ
40-49 years, male
366.5
GTEX-117YW-1526-SM-5EGGP
50-59 years, male
359.4
GTEX-XQ3S-2726-SM-4BOP2
20-29 years, male
357.5
GTEX-1C6VR-1526-SM-7MKG9
60-69 years, male
354.9
GTEX-13112-0226-SM-5P9IV
50-59 years, male
347.0
GTEX-S7PM-0626-SM-4AD4Q
60-69 years, male
346.1
GTEX-WK11-0326-SM-4OOS6
50-59 years, male
335.2
GTEX-QEG4-0126-SM-48TZE
30-39 years, male
334.6
GTEX-17MFQ-0826-SM-793C8
20-29 years, male
333.6
GTEX-11TUW-2226-SM-5EQL9
60-69 years, male
322.6
GTEX-1EKGG-2726-SM-79OOD
60-69 years, male
320.5
GTEX-11WQC-2326-SM-5EQKE
60-69 years, male
282.0
GTEX-1C64N-2826-SM-7IGP5
50-59 years, male
278.0
GTEX-11ZUS-2726-SM-5FQUA
30-39 years, male
254.2
GTEX-11NV4-1726-SM-5N9FC
60-69 years, male
250.8
GTEX-13FHP-2826-SM-5IJFW
50-59 years, male
200.9
GTEX-111VG-1926-SM-5GIDO
60-69 years, male
159.9
Show allShow less
TESTIS - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.