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On the top, protein expression in current human tissue, based on all annotated cell types, is reported with the units not detected (n), low (l), medium (m) and high (h). Underneath, protein expression in each annotated cell type are reported using the same units.
Protein expression data is based on knowledge-based annotation. For genes where more than one antibody has been used, a collective score is set.
If knowledge-based annotation could not be performed for a gene, no data is displayed here. View antibody staining data further down this page.
A summary of mRNA expression data available for current tissue based on several datasets. The mRNA expression levels in human tissues are based on RNA-seq data generated by the Human Protein Atlas (HPA), Genotype-Tissue Expression (GTEx) portal and CAGE data generated by the FANTOM5 consortium. Consensus normalized expression (NX) levels for human tissues was created by combining the data from these three transcriptomics datasets (HPA, GTEx and FANTOM5).
The mRNA expression levels in pig are based on RNA-seq data generated by the Human Protein Atlas (HPA), and for mouse, HPA data and in situ hybridization generated by the Allen brain atlas are reported.
Scroll down to view mRNA expression data in more detail.
The RNA-seq details section shows detailed information about the individual samples used for the transcript profiling and results of the RNA-seq analysis.
Information about each individual sample is listed below, including gender, age, a tissue section image and estimated fractions of cell types. pTPM (transcripts per million) values give a quantification of the gene abundance which is comparable between different genes and samples.
Cells in seminiferous ducts: 65 Other cell types: 35
Male, age 34
Testis sample 278 pTPM: 877.1
Cell types%
Cells in seminiferous ducts: 75 Other cell types: 25
Male, age 62
Testis sample 279 pTPM: 922.7
Cell types%
Cells in seminiferous ducts: 70 Other cell types: 30
Male, age 34
Testis sample 280 pTPM: 329.5
Cell types%
Cells in seminiferous ducts: 60 Other cell types: 40
Male, age 26
Testis sample 281 pTPM: 1110.1
Cell types%
Cells in seminiferous ducts: 55 Other cell types: 45
Male, age 50
Testis sample 282 pTPM: 976.1
Cell types%
Cells in seminiferous ducts: 85 Other cell types: 15
Male, age 37
Testis sample 283 pTPM: 740.2
Cell types%
Cells in seminiferous ducts: 35 Other cell types: 65
Male, age 51
Testis sample 349 pTPM: 244
Cell types%
Cells in seminiferous ducts: 65 Other cell types: 35
Male, age 34
Testis sample 354 pTPM: 546.1
Cell types%
Cells in seminiferous ducts: 70 Other cell types: 30
Male, age 26
Testis sample 371 pTPM: 583.5
Cell types%
Cells in seminiferous ducts: 75 Other cell types: 25
TESTIS - GTEx RNA-seqi
RNA-Seq data generated by the Genotype-Tissue Expression (GTEx) project from human tissues is reported as average pTPM. More information can be found on the GTEx portal.
Distribution across the dataset is visualized with box plots, shown as median and 25th and 75th percentiles. Points are displayed as outliers if they are above or below 1.5 times the interquartile range. pTPM values of the individual samples are presented next to the box plot.
Max pTPM: 2448.4
Min pTPM: 0.5
Std pTPM: 463.2
Median pTPM: 1071.3
GTEX-17HII-0126-SM-7KFSS
50-59 years, male
2448.4
GTEX-11NV4-1726-SM-5N9FC
60-69 years, male
2245.1
GTEX-1A3MX-1026-SM-731F8
60-69 years, male
2190.5
GTEX-11WQC-2326-SM-5EQKE
60-69 years, male
2048.0
GTEX-WK11-0326-SM-4OOS6
50-59 years, male
1920.6
GTEX-13VXU-0726-SM-5J2O7
50-59 years, male
1902.7
GTEX-15G19-1326-SM-6LPIR
60-69 years, male
1833.5
GTEX-1GMRU-0626-SM-7MKH2
50-59 years, male
1815.5
GTEX-17HHY-0726-SM-7IGM6
50-59 years, male
1794.0
GTEX-14C39-0526-SM-664OF
40-49 years, male
1754.2
GTEX-QEG4-0126-SM-48TZE
30-39 years, male
1730.0
GTEX-13FHP-2826-SM-5IJFW
50-59 years, male
1729.1
GTEX-139TS-1726-SM-5IJG5
60-69 years, male
1723.1
GTEX-13112-0226-SM-5P9IV
50-59 years, male
1716.1
GTEX-1B8SF-2926-SM-731DL
60-69 years, male
1693.6
GTEX-T6MN-2026-SM-4DM7L
50-59 years, male
1687.6
GTEX-13N1W-2626-SM-5IJEP
70-79 years, male
1673.1
GTEX-11ZUS-2726-SM-5FQUA
30-39 years, male
1670.1
GTEX-XQ3S-2726-SM-4BOP2
20-29 years, male
1666.0
GTEX-117YW-1526-SM-5EGGP
50-59 years, male
1654.5
GTEX-17KNJ-1126-SM-7KFT6
50-59 years, male
1647.2
GTEX-Z93S-1726-SM-5HL8G
50-59 years, male
1628.9
GTEX-PW2O-1426-SM-48TCD
20-29 years, male
1624.3
GTEX-17GQL-1726-SM-718B8
60-69 years, male
1605.9
GTEX-OOBK-2126-SM-3LK5T
40-49 years, male
1602.9
GTEX-1C4CL-1926-SM-731DY
30-39 years, male
1598.1
GTEX-12ZZY-0126-SM-5LZV2
60-69 years, male
1583.7
GTEX-WHSE-0426-SM-4M1XO
20-29 years, male
1573.9
GTEX-ZTSS-1526-SM-51MTC
40-49 years, male
1562.1
GTEX-1C64O-2126-SM-7IGPB
60-69 years, male
1558.2
GTEX-11EQ8-1426-SM-5EGJR
60-69 years, male
1544.5
GTEX-16YQH-1826-SM-6LPJW
30-39 years, male
1525.2
GTEX-18465-1526-SM-7KFTV
50-59 years, male
1525.1
GTEX-REY6-0126-SM-48FDT
60-69 years, male
1523.0
GTEX-13OVL-0426-SM-5IFG6
50-59 years, male
1521.2
GTEX-WYJK-1826-SM-4ONDM
50-59 years, male
1516.2
GTEX-14PJ2-2026-SM-6AJAQ
50-59 years, male
1511.0
GTEX-S7PM-0626-SM-4AD4Q
60-69 years, male
1506.6
GTEX-14ABY-0626-SM-5Q5C9
50-59 years, male
1501.3
GTEX-XLM4-1526-SM-4AT6D
60-69 years, male
1492.3
GTEX-ZVTK-0126-SM-57WDG
20-29 years, male
1490.0
GTEX-QV31-1126-SM-4R1K4
30-39 years, male
1489.9
GTEX-144GM-0426-SM-5Q5C8
20-29 years, male
1478.7
GTEX-1CB4G-2426-SM-7DUGM
60-69 years, male
1469.6
GTEX-18A7B-1526-SM-7KFTH
70-79 years, male
1463.8
GTEX-ZAB5-2426-SM-5CVMW
50-59 years, male
1446.1
GTEX-SNOS-1126-SM-4DM67
40-49 years, male
1439.2
GTEX-TKQ2-1526-SM-4DXUN
50-59 years, male
1424.4
GTEX-Q2AI-1226-SM-48U14
40-49 years, male
1422.3
GTEX-139TT-2226-SM-5LZWO
60-69 years, male
1405.4
GTEX-V1D1-2126-SM-4JBH4
50-59 years, male
1403.9
GTEX-1C64N-2826-SM-7IGP5
50-59 years, male
1402.6
GTEX-R55E-0726-SM-48FCZ
20-29 years, male
1397.2
GTEX-Y111-2426-SM-4TT23
50-59 years, male
1390.8
GTEX-1F6IF-0526-SM-7MKHD
60-69 years, male
1385.3
GTEX-1EU9M-1626-SM-7EWF2
60-69 years, male
1384.1
GTEX-111VG-1926-SM-5GIDO
60-69 years, male
1373.7
GTEX-12WSL-2326-SM-5DUXQ
50-59 years, male
1372.5
GTEX-S95S-1126-SM-4B64E
60-69 years, male
1372.2
GTEX-11GS4-2026-SM-5N9CP
60-69 years, male
1371.7
GTEX-U8T8-1126-SM-4DXUE
60-69 years, male
1367.9
GTEX-UPJH-0126-SM-4IHLL
50-59 years, male
1364.7
GTEX-OHPM-2126-SM-3LK75
50-59 years, male
1358.5
GTEX-131XE-0426-SM-5IJF4
50-59 years, male
1357.8
GTEX-18A6Q-2626-SM-7KFT3
60-69 years, male
1357.4
GTEX-ZQUD-2026-SM-51MSM
30-39 years, male
1353.2
GTEX-144GL-0726-SM-5LU4P
50-59 years, male
1342.6
GTEX-S4Q7-1226-SM-4AD5I
20-29 years, male
1337.3
GTEX-OIZH-2126-SM-3NB1P
50-59 years, male
1333.2
GTEX-15SHV-1926-SM-6LPI1
60-69 years, male
1323.9
GTEX-XGQ4-2026-SM-4AT6G
50-59 years, male
1320.1
GTEX-13NZB-2026-SM-5MR4M
30-39 years, male
1318.5
GTEX-ZA64-1626-SM-5CVME
20-29 years, male
1316.0
GTEX-183FY-1926-SM-7KFRI
20-29 years, male
1308.4
GTEX-1C6VR-1526-SM-7MKG9
60-69 years, male
1302.8
GTEX-11DXZ-2126-SM-59881
50-59 years, male
1298.8
GTEX-12WSH-0326-SM-5GCNH
50-59 years, male
1297.9
GTEX-ZVZP-2226-SM-57WBF
50-59 years, male
1295.8
GTEX-YF7O-2026-SM-4W1YE
50-59 years, male
1287.7
GTEX-XAJ8-1326-SM-47JYT
40-49 years, male
1286.2
GTEX-Y5V6-1726-SM-4VDSZ
60-69 years, male
1268.7
GTEX-V955-1926-SM-4KL1L
60-69 years, male
1267.9
GTEX-QEG5-0126-SM-4R1JR
20-29 years, male
1267.8
GTEX-1B996-2326-SM-731EC
60-69 years, male
1262.8
GTEX-1A3MV-0726-SM-72D5M
30-39 years, male
1259.6
GTEX-11O72-0726-SM-5P9GO
40-49 years, male
1241.2
GTEX-17MF6-2726-SM-7IGMZ
40-49 years, male
1237.0
GTEX-18D9B-2726-SM-72D73
60-69 years, male
1236.4
GTEX-11NUK-2626-SM-5A5MB
50-59 years, male
1226.5
GTEX-145MH-2326-SM-5O9AW
50-59 years, male
1226.2
GTEX-14E1K-1926-SM-73KWS
50-59 years, male
1224.7
GTEX-11OF3-1826-SM-5987N
60-69 years, male
1222.5
GTEX-OXRL-2126-SM-3NM98
50-59 years, male
1221.0
GTEX-12696-0226-SM-5EGL3
60-69 years, male
1215.9
GTEX-145MO-0126-SM-5S2QU
60-69 years, male
1209.7
GTEX-QLQW-1026-SM-447A9
30-39 years, male
1209.0
GTEX-111CU-1726-SM-5EGHM
50-59 years, male
1207.3
GTEX-13NZA-2526-SM-5IJFX
60-69 years, male
1206.1
GTEX-132QS-1226-SM-5P9GD
60-69 years, male
1203.7
GTEX-YEC3-1726-SM-5IFIK
50-59 years, male
1190.2
GTEX-ZT9X-1426-SM-5DUX1
40-49 years, male
1186.7
GTEX-13FLW-2126-SM-5N9FD
50-59 years, male
1178.0
GTEX-ZV7C-2026-SM-5NQ8F
50-59 years, male
1176.6
GTEX-14BIL-2226-SM-73KWF
50-59 years, male
1175.1
GTEX-OOBJ-2126-SM-3NB1N
60-69 years, male
1171.8
GTEX-1AX9J-1226-SM-72D6P
60-69 years, male
1169.3
GTEX-WZTO-0326-SM-4PQYZ
40-49 years, male
1168.1
GTEX-ZPU1-2126-SM-57WED
40-49 years, male
1167.8
GTEX-14C38-0126-SM-5YY9V
50-59 years, male
1165.9
GTEX-13OW5-2526-SM-5L3I1
60-69 years, male
1164.0
GTEX-ZYT6-2726-SM-5GICP
30-39 years, male
1163.4
GTEX-TKQ1-0926-SM-4DXU2
30-39 years, male
1161.3
GTEX-15RIF-1826-SM-6M469
20-29 years, male
1161.0
GTEX-1AX9I-1826-SM-72D5I
60-69 years, male
1158.0
GTEX-11NSD-1026-SM-5N9BE
20-29 years, male
1153.0
GTEX-16XZZ-0626-SM-6M47T
60-69 years, male
1148.4
GTEX-N7MS-0126-SM-3TW8O
60-69 years, male
1135.1
GTEX-S3XE-1526-SM-4AD5A
50-59 years, male
1133.7
GTEX-XBEC-0126-SM-4GIDT
50-59 years, male
1127.8
GTEX-X5EB-2026-SM-4E3KA
40-49 years, male
1120.5
GTEX-ZYFC-0126-SM-5GIEH
50-59 years, male
1118.1
GTEX-117XS-2026-SM-5GID1
60-69 years, male
1112.9
GTEX-S7SE-0326-SM-4AT5Q
50-59 years, male
1101.1
GTEX-T5JC-0726-SM-4DM55
20-29 years, male
1093.8
GTEX-X3Y1-2626-SM-4PQZI
40-49 years, male
1093.2
GTEX-U4B1-1526-SM-4DXSL
40-49 years, male
1085.8
GTEX-18A66-0726-SM-72D77
60-69 years, male
1081.6
GTEX-13111-1526-SM-5EGJX
50-59 years, male
1079.2
GTEX-1EKGG-2726-SM-79OOD
60-69 years, male
1071.7
GTEX-12WSI-2126-SM-5GCMV
50-59 years, male
1071.3
GTEX-WHSB-2126-SM-4M1XF
50-59 years, male
1068.0
GTEX-Q2AH-1526-SM-48TZG
40-49 years, male
1059.1
GTEX-1212Z-0326-SM-5FQSJ
60-69 years, male
1046.3
GTEX-WH7G-1926-SM-4LVMM
40-49 years, male
1038.7
GTEX-13O1R-0726-SM-5IJEI
60-69 years, male
1038.1
GTEX-111FC-1926-SM-5GZYC
60-69 years, male
1032.7
GTEX-YEC4-1526-SM-4W1YU
40-49 years, male
1021.8
GTEX-16Z82-2726-SM-7KULV
60-69 years, male
1021.1
GTEX-13N2G-0126-SM-5N9DV
60-69 years, male
1021.0
GTEX-ZZ64-1126-SM-5GZXY
20-29 years, male
1019.6
GTEX-SNMC-1026-SM-4DM7K
20-29 years, male
1017.8
GTEX-14E6E-1026-SM-664N9
30-39 years, male
1016.1
GTEX-YJ89-0626-SM-4TT3Z
60-69 years, male
1012.1
GTEX-14PJ3-1626-SM-664O5
50-59 years, male
1007.0
GTEX-16AAH-1626-SM-7EWE2
50-59 years, male
999.3
GTEX-T8EM-0126-SM-4DM5R
40-49 years, male
988.9
GTEX-12C56-1426-SM-5FQSW
20-29 years, male
987.8
GTEX-1B8KE-1426-SM-7EWEL
50-59 years, male
984.9
GTEX-14DAR-1126-SM-793AT
50-59 years, male
983.6
GTEX-11EQ9-1926-SM-5PNVV
30-39 years, male
981.4
GTEX-144GN-1626-SM-5Q5BU
50-59 years, male
978.0
GTEX-X261-2326-SM-4PQYU
50-59 years, male
975.6
GTEX-111YS-2026-SM-5EGGL
60-69 years, male
973.3
GTEX-U3ZM-1626-SM-4DXSK
40-49 years, male
972.0
GTEX-14753-0626-SM-5Q5CY
60-69 years, male
971.0
GTEX-ZDYS-1326-SM-5IJFF
50-59 years, male
969.1
GTEX-WOFM-1126-SM-4OOSB
20-29 years, male
968.6
GTEX-13NYB-2226-SM-5MR58
40-49 years, male
964.4
GTEX-P4QS-2126-SM-3NMCF
60-69 years, male
962.5
GTEX-XBED-2026-SM-4AT5D
60-69 years, male
960.7
GTEX-13O61-2026-SM-5J2M6
60-69 years, male
946.6
GTEX-17HHE-1226-SM-793C6
50-59 years, male
945.2
GTEX-XMK1-2026-SM-4B65K
40-49 years, male
942.1
GTEX-ZT9W-2226-SM-57WFU
50-59 years, male
942.1
GTEX-11EI6-2226-SM-5EGJM
60-69 years, male
930.6
GTEX-18QFQ-1926-SM-72D5H
30-39 years, male
930.1
GTEX-WVLH-2626-SM-4MVNV
50-59 years, male
926.9
GTEX-13O21-1226-SM-5J2MK
50-59 years, male
908.3
GTEX-11TUW-2226-SM-5EQL9
60-69 years, male
902.7
GTEX-11LCK-2326-SM-5HL53
30-39 years, male
897.4
GTEX-1E1VI-1026-SM-7MKGQ
50-59 years, male
891.6
GTEX-11P82-1526-SM-5BC5M
20-29 years, male
891.4
GTEX-RUSQ-2126-SM-47JXK
50-59 years, male
889.3
GTEX-WFG8-1926-SM-4LVM1
20-29 years, male
888.1
GTEX-ZUA1-2726-SM-59HLJ
40-49 years, male
883.0
GTEX-QMRM-1526-SM-4R1K6
50-59 years, male
877.7
GTEX-RM2N-1326-SM-48FCW
50-59 years, male
866.8
GTEX-139T6-1226-SM-5IFFC
50-59 years, male
864.5
GTEX-15RIE-0626-SM-6M47G
60-69 years, male
863.4
GTEX-13OVH-0726-SM-5N9BU
70-79 years, male
860.4
GTEX-16MTA-1126-SM-6LPJA
50-59 years, male
853.3
GTEX-NFK9-0126-SM-3LK5H
40-49 years, male
842.5
GTEX-R55D-0126-SM-48FEL
50-59 years, male
841.0
GTEX-U8XE-0126-SM-4E3I3
30-39 years, male
839.2
GTEX-R55C-1426-SM-48FED
40-49 years, male
814.4
GTEX-17MFQ-0826-SM-793C8
20-29 years, male
809.6
GTEX-14E7W-0726-SM-664OK
70-79 years, male
805.1
GTEX-17F9E-1626-SM-7EWDB
50-59 years, male
802.3
GTEX-QV44-1726-SM-4R1KG
50-59 years, male
800.5
GTEX-1BAJH-0726-SM-7IGMF
70-79 years, male
797.7
GTEX-1AX9K-1826-SM-731CY
50-59 years, male
788.0
GTEX-147GR-0626-SM-5S2PK
60-69 years, male
783.0
GTEX-S33H-0126-SM-4AD62
30-39 years, male
781.2
GTEX-17HGU-0726-SM-7DUFN
50-59 years, male
774.9
GTEX-14C5O-2326-SM-73KYU
60-69 years, male
745.4
GTEX-14A5H-0626-SM-5TDCO
60-69 years, male
739.5
GTEX-11TT1-2226-SM-5GU6B
20-29 years, male
733.8
GTEX-1399R-1626-SM-5P9GG
30-39 years, male
728.3
GTEX-YFCO-1726-SM-4W21S
40-49 years, male
728.2
GTEX-RN64-2326-SM-48FDW
50-59 years, male
718.9
GTEX-VJYA-1426-SM-4KL1Y
60-69 years, male
711.1
GTEX-ZTX8-1126-SM-51MRM
20-29 years, male
704.7
GTEX-14B4R-1026-SM-5TDDS
30-39 years, male
696.6
GTEX-13OVK-2226-SM-6LPJY
40-49 years, male
693.2
GTEX-145LT-0426-SM-5LUAP
40-49 years, male
691.4
GTEX-11P7K-1026-SM-5HL6Y
30-39 years, male
687.3
GTEX-12WSM-1326-SM-5GCP9
60-69 years, male
681.4
GTEX-O5YT-2126-SM-3MJGD
20-29 years, male
670.8
GTEX-OIZI-0126-SM-3NB13
40-49 years, male
657.7
GTEX-147JS-0126-SM-5S2TW
60-69 years, male
657.1
GTEX-1339X-1926-SM-5PNVP
40-49 years, male
643.6
GTEX-XPT6-1626-SM-4B655
20-29 years, male
631.5
GTEX-15RJ7-1826-SM-7KUMJ
40-49 years, male
626.2
GTEX-RWSA-2426-SM-47JXR
40-49 years, male
621.6
GTEX-14A6H-2326-SM-5Q5B5
30-39 years, male
605.6
GTEX-QLQ7-1426-SM-4R1JX
60-69 years, male
604.0
GTEX-14PHX-1426-SM-69LPP
50-59 years, male
592.8
GTEX-YB5E-1926-SM-5IFIG
40-49 years, male
586.1
GTEX-14PJN-1626-SM-68727
30-39 years, male
581.4
GTEX-13FTW-1326-SM-5LZZD
40-49 years, male
559.7
GTEX-14PJ4-1726-SM-664OC
60-69 years, male
549.9
GTEX-U3ZH-1526-SM-4DXV1
30-39 years, male
546.9
GTEX-15SKB-0326-SM-6M477
60-69 years, male
541.9
GTEX-147F4-0626-SM-5LUAK
50-59 years, male
484.6
GTEX-13OW6-0126-SM-5IJGM
50-59 years, male
481.3
GTEX-PLZ5-1526-SM-3P5ZX
50-59 years, male
471.7
GTEX-1CAMR-1526-SM-79OLP
20-29 years, male
452.7
GTEX-1CB4I-0926-SM-7MKFY
50-59 years, male
446.4
GTEX-1399T-1526-SM-5P9J6
60-69 years, male
444.7
GTEX-XPVG-2226-SM-4B65U
50-59 years, male
434.1
GTEX-Y8E4-2226-SM-5LU94
40-49 years, male
393.8
GTEX-Y3I4-2026-SM-4TT6Z
50-59 years, male
384.8
GTEX-11ONC-2226-SM-5HL6D
60-69 years, male
381.4
GTEX-WFON-2026-SM-4LVMW
40-49 years, male
377.7
GTEX-ZDTT-2026-SM-5K7TY
60-69 years, male
377.1
GTEX-PLZ6-1226-SM-3P5ZS
30-39 years, male
320.6
GTEX-ZLFU-2026-SM-4WWG2
40-49 years, male
293.3
GTEX-1AYCT-1026-SM-79ONR
50-59 years, male
282.9
GTEX-ZAB4-0126-SM-5CVMG
40-49 years, male
280.8
GTEX-145MF-1726-SM-5LU9H
50-59 years, male
274.5
GTEX-Y9LG-1726-SM-4VBQE
30-39 years, male
265.3
GTEX-1E2YA-2826-SM-7EPIN
50-59 years, male
264.4
GTEX-18A67-1826-SM-7KFT7
50-59 years, male
237.9
GTEX-12BJ1-1326-SM-5BC5P
60-69 years, male
140.7
GTEX-11DXY-0226-SM-5H123
60-69 years, male
117.6
GTEX-13OW8-0526-SM-5KM24
60-69 years, male
114.3
GTEX-117YX-2026-SM-5GIEF
50-59 years, male
100.5
GTEX-13QJ3-0226-SM-5S2PU
50-59 years, male
73.3
GTEX-1C6VS-1126-SM-7EWEW
60-69 years, male
48.0
GTEX-1B8KZ-1826-SM-73KVJ
50-59 years, male
43.2
GTEX-S4Z8-2126-SM-4AD5H
60-69 years, male
42.7
GTEX-1AX8Z-2826-SM-73KTZ
60-69 years, male
27.6
GTEX-WY7C-2226-SM-4ONCS
50-59 years, male
18.1
GTEX-1C6VQ-1426-SM-73KTW
50-59 years, male
6.6
GTEX-NPJ8-1226-SM-3MJHM
40-49 years, male
4.5
GTEX-17F97-2726-SM-7IGQ4
50-59 years, male
3.7
GTEX-148VJ-2526-SM-5TDEI
70-79 years, male
3.3
GTEX-1CB4J-1626-SM-79OOK
60-69 years, male
0.5
GTEX-WVJS-2726-SM-4MVNU
50-59 years, male
0.5
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TESTIS - FANTOM5 CAGEi
RNA expression in human tissues obtained through Cap Analysis of Gene Expression (CAGE) generated by the FANTOM5 project are reported as Scaled Tags Per Million.